Ant - Wikipedia
Jump to content
Main menu
Main menu
move to sidebar
hide
Navigation
Main pageContentsCurrent eventsRandom articleAbout WikipediaContact usDonate
Contribute
HelpLearn to editCommunity portalRecent changesUpload file
Search
Search
Create account
Log in
Personal tools
Create account Log in
Pages for logged out editors learn more
ContributionsTalk
Contents
move to sidebar
hide
(Top)
1Etymology
2Taxonomy and evolution
3Distribution and diversity
4Morphology
Toggle Morphology subsection
4.1Head
4.2Mesosoma
4.3Metasoma
4.4Polymorphism
4.5Genome size
5Life cycle
Toggle Life cycle subsection
5.1Reproduction
5.2Nests, colonies, and supercolonies
6Behaviour and ecology
Toggle Behaviour and ecology subsection
6.1Communication
6.2Defence
6.3Learning
6.4Nest construction
6.5Cultivation of food
6.6Navigation
6.7Locomotion
6.8Cooperation and competition
6.9Relationships with other organisms
7Relationship with humans
Toggle Relationship with humans subsection
7.1As food
7.2As pests
7.3In science and technology
7.4As pets
7.5In culture
8See also
9References
Toggle References subsection
9.1Cited texts
10Further reading
11External links
Toggle the table of contents
Ant
193 languages
AfrikaansAlemannischአማርኛअंगिकाÆngliscالعربيةAragonésܐܪܡܝܐԱրեւմտահայերէնArmãneashtiঅসমীয়াAsturianuAtikamekwAvañe'ẽAymar aruAzərbaycancaتۆرکجهBasa BaliBamanankanবাংলাBanjarBân-lâm-gúBasa BanyumasanБашҡортсаБеларускаяБеларуская (тарашкевіца)Bikol CentralБългарскиབོད་ཡིགBosanskiBrezhonegБуряадCatalàЧӑвашлаCebuanoČeštinaChiShonaCymraegDagbanliDanskDeutschDiné bizaadEestiΕλληνικάEmiliàn e rumagnòlEspañolEsperantoEuskaraفارسیFiji HindiFrançaisGaeilgeGàidhligGalegoГӀалгӀайગુજરાતી客家語/Hak-kâ-ngîХальмг한국어Հայերենहिन्दीHrvatskiIdoIlokanoBahasa IndonesiaInterlinguaИронÍslenskaItalianoעבריתJawaKabɩyɛಕನ್ನಡKapampanganქართულიकॉशुर / کٲشُرҚазақшаKernowekKiswahiliКомиKongoKreyòl ayisyenKurdîКыргызчаКырык марыLadinЛаккуLatinaLatviešuLëtzebuergeschЛезгиLietuviųLi NihaLimburgsLivvinkarjalaLombardMagyarМакедонскиMalagasyമലയാളംमराठीმარგალურიمصرىمازِرونیBahasa Melayuꯃꯤꯇꯩ ꯂꯣꯟ閩東語 / Mìng-dĕ̤ng-ngṳ̄Монголမြန်မာဘာသာNa Vosa VakavitiNederlandsNedersaksiesनेपालीनेपाल भाषा日本語НохчийнNordfriiskNorsk bokmålNorsk nynorskOccitanОлык марийଓଡ଼ିଆOʻzbekcha / ўзбекчаਪੰਜਾਬੀPälzischPangcahپنجابیپښتوПерем комиPinayuananPlattdüütschPolskiPortuguêsRipoarischRomânăRumantschRuna SimiРусиньскыйРусскийСаха тылаSakizayaᱥᱟᱱᱛᱟᱲᱤSarduScotsShqipSicilianuසිංහලSimple EnglishسنڌيSlovenčinaSlovenščinaŚlůnskiSoomaaligaکوردیСрпски / srpskiSrpskohrvatski / српскохрватскиSundaSuomiSvenskaTagalogதமிழ்TaclḥitTaqbaylitТатарча / tatarçaతెలుగుไทยThuɔŋjäŋТоҷикӣᏣᎳᎩತುಳುTürkçeTürkmençeTyapУдмуртУкраїнськаاردوئۇيغۇرچە / UyghurcheVahcuenghVepsän kel’Tiếng ViệtWalonWayuunaiki文言West-VlamsWinarayWolof吴语ייִדיש粵語ZazakiŽemaitėška中文Batak Toba
Edit links
ArticleTalk
English
ReadView sourceView history
Tools
Tools
move to sidebar
hide
Actions
ReadView sourceView history
General
What links hereRelated changesUpload fileSpecial pagesPermanent linkPage informationCite this pageGet shortened URLDownload QR codeWikidata item
Print/export
Download as PDFPrintable version
In other projects
Wikimedia CommonsWikispeciesWikinewsWikiquoteWikiversity
From Wikipedia, the free encyclopedia
Family of insects
For other uses, see Ant (disambiguation).
AntsTemporal range: 100–0 Ma[1]
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N
Late Albian – Present
Fire ants
Scientific classification
Domain:
Eukaryota
Kingdom:
Animalia
Phylum:
Arthropoda
Class:
Insecta
Order:
Hymenoptera
Infraorder:
Aculeata
Superfamily:
Formicoidea
Family:
FormicidaeLatreille, 1809
Type species
Formica rufaLinnaeus, 1761
Subfamilies
Agroecomyrmecinae
Amblyoponinae (incl. "Apomyrminae")
Aneuretinae
†Brownimeciinae
Dolichoderinae
Dorylinae
Ectatomminae
†Formiciinae
Formicinae
†Haidomyrmecinae
Heteroponerinae
Leptanillinae
Martialinae
Myrmeciinae (incl. "Nothomyrmeciinae")
Myrmicinae
Paraponerinae
Ponerinae
Proceratiinae
Pseudomyrmecinae
†Sphecomyrminae
†Zigrasimeciinae
Cladogram of subfamilies
Formicoid
Myrmicinae
Ectatomminae
Heteroponerinae
Formicinae
Dolichoderinae
Aneuretinae
Pseudomyrmecinae
Myrmeciinae
Dorylinae‡
Poneroid
Ponerinae
Agroecomyrmecinae
Paraponerinae
Proceratiinae
Amblyoponinae
Apomyrminae
Leptanillinae
Martialinae
A phylogeny of the extant ant subfamilies.[2][3][4]
*Cerapachyinae is paraphyletic
‡ The previous dorylomorph subfamilies - Ecitoninae, Aenictinae, Aenictogitoninae, Cerapachyinae, Leptanilloidinae - were synonymized under Dorylinae by Brady et al. in 2014[5]
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists.
Ants form colonies that range in size from a few dozen predatory individuals living in small natural cavities to highly organised colonies that may occupy large territories and consist of millions of individuals. Larger colonies consist of various castes of sterile, wingless females, most of which are workers (ergates), as well as soldiers (dinergates) and other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called "queens" (gynes). The colonies are described as superorganisms because the ants appear to operate as a unified entity, collectively working together to support the colony.
Ants gathering food (video)
Ants have colonised almost every landmass on Earth. The only places lacking indigenous ants are Antarctica and a few remote or inhospitable islands. Ants thrive in moist tropical ecosystems and may exceed the combined biomass of wild birds and mammals. Their success in so many environments has been attributed to their social organisation and their ability to modify habitats, tap resources, and defend themselves. Their long co-evolution with other species has led to mimetic, commensal, parasitic, and mutualistic relationships.
Ant societies have division of labour, communication between individuals, and an ability to solve complex problems. These parallels with human societies have long been an inspiration and subject of study. Many human cultures make use of ants in cuisine, medication, and rites. Some species are valued in their role as biological pest control agents. Their ability to exploit resources may bring ants into conflict with humans, however, as they can damage crops and invade buildings. Some species, such as the red imported fire ant (Solenopsis invicta) of South America, are regarded as invasive species in other parts of the world, establishing themselves in areas where they have been introduced accidentally.
Etymology
The word ant and the archaic word emmet[6] are derived from ante, emete of Middle English, which come from ǣmette of Old English; these are all related to Low Saxon e(e)mt, empe and varieties (Old Saxon emeta) and to German Ameise (Old High German āmeiza). All of these words come from West Germanic *ǣmaitjōn, and the original meaning of the word was "the biter" (from Proto-Germanic *ai-, "off, away" + *mait- "cut").[7][8]
The family name Formicidae is derived from the Latin formīca ("ant")[9] from which the words in other Romance languages, such as the Portuguese formiga, Italian formica, Spanish hormiga, Romanian furnică, and French fourmi are derived. It has been hypothesised that a Proto-Indo-European word *morwi- was the root for Sanskrit vamrah, Greek μύρμηξ mýrmēx, Old Church Slavonic mraviji, Old Irish moirb, Old Norse maurr, Dutch mier, Swedish myra, Danish myre, Middle Dutch miere, and Crimean Gothic miera.[10][11]
Taxonomy and evolution
Aculeata
Chrysidoidea
Vespidae
Rhopalosomatidae
Pompilidae
Tiphiidae
Scolioidea
Apoidea
Formicidae
Phylogenetic position of the Formicidae[12]
The family Formicidae belongs to the order Hymenoptera, which also includes sawflies, bees, and wasps. Ants evolved from a lineage within the stinging wasps, and a 2013 study suggests that they are a sister group of the Apoidea.[12] In 1966, E. O. Wilson and his colleagues identified the fossil remains of an ant (Sphecomyrma) that lived in the Cretaceous period. The specimen, trapped in amber dating back to around 92 million years ago, has features found in some wasps, but not found in modern ants.[13] The oldest fossils of ants date to the mid-Cretaceous, around 100 million years ago, which belong to extinct stem-groups such as the Haidomyrmecinae, Sphecomyrminae and Zigrasimeciinae, with modern ant subfamilies appearing towards the end of the Cretaceous around 80–70 million years ago.[14] Ants diversified and assumed ecological dominance around 60 million years ago.[15][1][16][17] Some groups, such as the Leptanillinae and Martialinae, are suggested to have diversified from early primitive ants that were likely to have been predators underneath the surface of the soil.[4][18]
Ants fossilised in Baltic amber
During the Cretaceous period, a few species of primitive ants ranged widely on the Laurasian supercontinent (the Northern Hemisphere). Their representation in the fossil record is poor, in comparison to the populations of other insects, representing only about 1% of fossil evidence of insects in the era. Ants became dominant after adaptive radiation at the beginning of the Paleogene period. By the Oligocene and Miocene, ants had come to represent 20–40% of all insects found in major fossil deposits. Of the species that lived in the Eocene epoch, around one in 10 genera survive to the present. Genera surviving today comprise 56% of the genera in Baltic amber fossils (early Oligocene), and 92% of the genera in Dominican amber fossils (apparently early Miocene).[15][19]
Termites live in colonies and are sometimes called "white ants", but termites are only distantly related to ants. They are the sub-order Isoptera, and together with cockroaches, they form the order Blattodea. Blattodeans are related to mantids, crickets, and other winged insects that do not undergo complete metamorphosis. Like ants, termites are eusocial, with sterile workers, but they differ greatly in the genetics of reproduction. The similarity of their social structure to that of ants is attributed to convergent evolution.[20] Velvet ants look like large ants, but are wingless female wasps.[21][22]
Distribution and diversity
Region
Number ofspecies [23]
Neotropics
2,162
Nearctic
580
Europe
180
Africa
2,500
Asia
2,080
Melanesia
275
Australia
985
Polynesia
42
Ants have a cosmopolitan distribution. They are found on all continents except Antarctica, and only a few large islands, such as Greenland, Iceland, parts of Polynesia and the Hawaiian Islands lack native ant species.[24][25] Ants occupy a wide range of ecological niches and exploit many different food resources as direct or indirect herbivores, predators and scavengers. Most ant species are omnivorous generalists, but a few are specialist feeders. There is considerable variation in ant abundance across habitats, peaking in the moist tropics to nearly six times that found in less suitable habitats.[26] Their ecological dominance has been examined primarily using estimates of their biomass: myrmecologist E. O. Wilson had estimated in 2009 that at any one time the total number of ants was between one and ten quadrillion (short scale) (i.e., between 1015 and 1016) and using this estimate he had suggested that the total biomass of all the ants in the world was approximately equal to the total biomass of the entire human race.[27] More careful estimates made in 2022 which take into account regional variations puts the global ant contribution at 12 megatons of dry carbon, which is about 20% of the total human contribution, but greater than that of the wild birds and mammals combined. This study also puts a conservative estimate of the ants at about 20 × 1015 (20 quadrillion).[28][29][30]
Ants range in size from 0.75 to 52 millimetres (0.030–2.0 in),[31][32] the largest species being the fossil Titanomyrma giganteum, the queen of which was 6 cm (2+1⁄2 in) long with a wingspan of 15 cm (6 in).[33] Ants vary in colour; most ants are yellow to red or brown to black, but a few species are green and some tropical species have a metallic lustre. More than 13,800 species are currently known[34] (with upper estimates of the potential existence of about 22,000; see the article List of ant genera), with the greatest diversity in the tropics. Taxonomic studies continue to resolve the classification and systematics of ants. Online databases of ant species, including AntWeb and the Hymenoptera Name Server, help to keep track of the known and newly described species.[34] The relative ease with which ants may be sampled and studied in ecosystems has made them useful as indicator species in biodiversity studies.[35][36]
Morphology
Diagram of a worker ant (Neoponera verenae)
Ants are distinct in their morphology from other insects in having geniculate (elbowed) antennae, metapleural glands, and a strong constriction of their second abdominal segment into a node-like petiole. The head, mesosoma, and metasoma are the three distinct body segments (formally tagmata). The petiole forms a narrow waist between their mesosoma (thorax plus the first abdominal segment, which is fused to it) and gaster (abdomen less the abdominal segments in the petiole). The petiole may be formed by one or two nodes (the second alone, or the second and third abdominal segments).[37] Tergosternal fusion, when the tergite and sternite of a segment fuse together, can occur partly or fully on the second, third and fourth abdominal segment and is used in identification. Fourth abdominal tergosternal fusion was formerly used as character that defined the poneromorph subfamilies, Ponerinae and relatives within their clade, but this is no longer considered a synapomorphic character.[38]
Like other arthropods, ants have an exoskeleton, an external covering that provides a protective casing around the body and a point of attachment for muscles, in contrast to the internal skeletons of humans and other vertebrates. Insects do not have lungs; oxygen and other gases, such as carbon dioxide, pass through their exoskeleton via tiny valves called spiracles. Insects also lack closed blood vessels; instead, they have a long, thin, perforated tube along the top of the body (called the "dorsal aorta") that functions like a heart, and pumps haemolymph toward the head, thus driving the circulation of the internal fluids. The nervous system consists of a ventral nerve cord that runs the length of the body, with several ganglia and branches along the way reaching into the extremities of the appendages.[39]
Head
Bull ant showing the powerful mandibles and the relatively large compound eyes that provide excellent vision
Ant head
An ant's head contains many sensory organs. Like most insects, ants have compound eyes made from numerous tiny lenses attached together. Ant eyes are good for acute movement detection, but do not offer a high resolution image. They also have three small ocelli (simple eyes) on the top of the head that detect light levels and polarization.[40] Compared to vertebrates, ants tend to have blurrier eyesight, particularly in smaller species,[41] and a few subterranean taxa are completely blind.[3] However, some ants, such as Australia's bulldog ant, have excellent vision and are capable of discriminating the distance and size of objects moving nearly a meter away.[42]
Two antennae ("feelers") are attached to the head; these organs detect chemicals, air currents, and vibrations; they also are used to transmit and receive signals through touch. The head has two strong jaws, the mandibles, used to carry food, manipulate objects, construct nests, and for defence.[39] In some species, a small pocket (infrabuccal chamber) inside the mouth stores food, so it may be passed to other ants or their larvae.[43]
Mesosoma
Both the legs and wings of the ant are attached to the mesosoma ("thorax"). The legs terminate in a hooked claw which allows them to hook on and climb surfaces.[44] Only reproductive ants (queens and males) have wings. Queens shed their wings after the nuptial flight, leaving visible stubs, a distinguishing feature of queens. In a few species, wingless queens (ergatoids) and males occur.[39]
Metasoma
The metasoma (the "abdomen") of the ant houses important internal organs, including those of the reproductive, respiratory (tracheae), and excretory systems. Workers of many species have their egg-laying structures modified into stings that are used for subduing prey and defending their nests.[39]
Polymorphism
Seven leafcutter ant workers of various castes (left) and two queens (right)
In the colonies of a few ant species, there are physical castes—workers in distinct size-classes, called minor, median, and major ergates. Often, the larger ants have disproportionately larger heads, and correspondingly stronger mandibles. These are known as macrergates while smaller workers are known as micrergates.[45] Although formally known as dinergates, such individuals are sometimes called "soldier" ants because their stronger mandibles make them more effective in fighting, although they still are workers and their "duties" typically do not vary greatly from the minor or median workers. In a few species, the median workers are absent, creating a sharp divide between the minors and majors.[46] Weaver ants, for example, have a distinct bimodal size distribution.[47][48] Some other species show continuous variation in the size of workers. The smallest and largest workers in Carebara diversa show nearly a 500-fold difference in their dry weights.[49]
Workers cannot mate; however, because of the haplodiploid sex-determination system in ants, workers of a number of species can lay unfertilised eggs that become fully fertile, haploid males. The role of workers may change with their age and in some species, such as honeypot ants, young workers are fed until their gasters are distended, and act as living food storage vessels. These food storage workers are called repletes.[50] For instance, these replete workers develop in the North American honeypot ant Myrmecocystus mexicanus. Usually the largest workers in the colony develop into repletes; and, if repletes are removed from the colony, other workers become repletes, demonstrating the flexibility of this particular polymorphism.[51] This polymorphism in morphology and behaviour of workers initially was thought to be determined by environmental factors such as nutrition and hormones that led to different developmental paths; however, genetic differences between worker castes have been noted in Acromyrmex sp.[52] These polymorphisms are caused by relatively small genetic changes; differences in a single gene of Solenopsis invicta can decide whether the colony will have single or multiple queens.[53] The Australian jack jumper ant (Myrmecia pilosula) has only a single pair of chromosomes (with the males having just one chromosome as they are haploid), the lowest number known for any animal, making it an interesting subject for studies in the genetics and developmental biology of social insects.[54][55]
Genome size
Genome size is a fundamental characteristic of an organism. Ants have been found to have tiny genomes, with the evolution of genome size suggested to occur through loss and accumulation of non-coding regions, mainly transposable elements, and occasionally by whole genome duplication.[56] This may be related to colonisation processes, but further studies are needed to verify this.[56]
Life cycle
Meat eater ant nest during swarming
The life of an ant starts from an egg; if the egg is fertilised, the progeny will be female diploid, if not, it will be male haploid. Ants develop by complete metamorphosis with the larva stages passing through a pupal stage before emerging as an adult. The larva is largely immobile and is fed and cared for by workers. Food is given to the larvae by trophallaxis, a process in which an ant regurgitates liquid food held in its crop. This is also how adults share food, stored in the "social stomach". Larvae, especially in the later stages, may also be provided solid food, such as trophic eggs, pieces of prey, and seeds brought by workers.[57]
The larvae grow through a series of four or five moults and enter the pupal stage. The pupa has the appendages free and not fused to the body as in a butterfly pupa.[58] The differentiation into queens and workers (which are both female), and different castes of workers, is influenced in some species by the nutrition the larvae obtain. Genetic influences and the control of gene expression by the developmental environment are complex and the determination of caste continues to be a subject of research.[59] Winged male ants, called drones (termed "aner" in old literature[60]), emerge from pupae along with the usually winged breeding females. Some species, such as army ants, have wingless queens. Larvae and pupae need to be kept at fairly constant temperatures to ensure proper development, and so often are moved around among the various brood chambers within the colony.[61]
A new ergate spends the first few days of its adult life caring for the queen and young. She then graduates to digging and other nest work, and later to defending the nest and foraging. These changes are sometimes fairly sudden, and define what are called temporal castes. Such age-based task-specialization or polyethism has been suggested as having evolved due to the high casualties involved in foraging and defence, making it an acceptable risk only for ants who are older and likely to die sooner from natural causes.[62][63] In the Brazilian ant Forelius pusillus, the nest entrance is closed from the outside to protect the colony from predatory ant species at sunset each day. About one to eight workers seal the nest entrance from the outside and they have no chance of returning to the nest and are in effect sacrificed.[64] Whether these seemingly suicidal workers are older workers has not been determined.[65]
Ant colonies can be long-lived. The queens can live for up to 30 years, and workers live from 1 to 3 years. Males, however, are more transitory, being quite short-lived and surviving for only a few weeks.[66] Ant queens are estimated to live 100 times as long as solitary insects of a similar size.[67]
Ants are active all year long in the tropics; however, in cooler regions, they survive the winter in a state of dormancy known as hibernation. The forms of inactivity are varied and some temperate species have larvae going into the inactive state (diapause), while in others, the adults alone pass the winter in a state of reduced activity.[68]
Alate male ant, Prenolepis imparis
Reproduction
Honey ants (Prenolepis imparis) mating
A wide range of reproductive strategies have been noted in ant species. Females of many species are known to be capable of reproducing asexually through thelytokous parthenogenesis.[69] Secretions from the male accessory glands in some species can plug the female genital opening and prevent females from re-mating.[70] Most ant species have a system in which only the queen and breeding females have the ability to mate. Contrary to popular belief, some ant nests have multiple queens, while others may exist without queens. Workers with the ability to reproduce are called "gamergates" and colonies that lack queens are then called gamergate colonies; colonies with queens are said to be queen-right.[71]
Drones can also mate with existing queens by entering a foreign colony, such as in army ants. When the drone is initially attacked by the workers, it releases a mating pheromone. If recognized as a mate, it will be carried to the queen to mate.[72] Males may also patrol the nest and fight others by grabbing them with their mandibles, piercing their exoskeleton and then marking them with a pheromone. The marked male is interpreted as an invader by worker ants and is killed.[73]
Most ants are univoltine, producing a new generation each year.[74] During the species-specific breeding period, winged females and winged males, known to entomologists as alates, leave the colony in what is called a nuptial flight. The nuptial flight usually takes place in the late spring or early summer when the weather is hot and humid. Heat makes flying easier and freshly fallen rain makes the ground softer for mated queens to dig nests.[75] Males typically take flight before the females. Males then use visual cues to find a common mating ground, for example, a landmark such as a pine tree to which other males in the area converge. Males secrete a mating pheromone that females follow. Males will mount females in the air, but the actual mating process usually takes place on the ground. Females of some species mate with just one male but in others they may mate with as many as ten or more different males, storing the sperm in their spermathecae.[76] In Cardiocondyla elegans, workers may transport newly emerged queens to other conspecific nests where wingless males from unrelated colonies can mate with them, a behavioural adaptation that may reduce the chances of inbreeding.[77]
Fertilised meat-eater ant queen beginning to dig a new colony
Mated females then seek a suitable place to begin a colony. There, they break off their wings using their tibial spurs and begin to lay and care for eggs. The females can selectively fertilise future eggs with the sperm stored to produce diploid workers or lay unfertilized haploid eggs to produce drones. The first workers to hatch, known as nanitics,[78] are weaker and smaller than later workers but they begin to serve the colony immediately. They enlarge the nest, forage for food, and care for the other eggs. Species that have multiple queens may have a queen leaving the nest along with some workers to found a colony at a new site,[76] a process akin to swarming in honeybees.
Nests, colonies, and supercolonies
The typical ant species has a colony occupying a single nest, housing one or more queens, where the brood is raised. There are however more than 150 species of ants in 49 genera that are known to have colonies consisting of multiple spatially separated nests. These polydomous (as opposed to monodomous) colonies have food and workers moving between the nests.[79] Membership to a colony is identified by the response of worker ants which identify whether another individual belongs to their own colony or not. A signature cocktail of body surface chemicals (also known as cuticular hydrocarbons or CHCs) forms the so-called colony odor which other members can recognize.[80] Some ant species appear to be less discriminating and in the Argentine ant Linepithema humile, workers carried from a colony anywhere in the southern US and Mexico are acceptable within other colonies in the same region. Similarly workers from colonies that have established in Europe are accepted by any other colonies within Europe but not by the colonies in the Americas. The interpretation of these observations has been debated and some have been termed these large populations as supercolonies[81][82] while others have termed the populations as unicolonial.[83]
Behaviour and ecology
Communication
Two Camponotus sericeus workers communicating through touch and pheromones
Ants communicate with each other using pheromones, sounds, and touch.[84] Since most ants live on the ground, they use the soil surface to leave pheromone trails that may be followed by other ants. In species that forage in groups, a forager that finds food marks a trail on the way back to the colony; this trail is followed by other ants, these ants then reinforce the trail when they head back with food to the colony. When the food source is exhausted, no new trails are marked by returning ants and the scent slowly dissipates. This behaviour helps ants deal with changes in their environment. For instance, when an established path to a food source is blocked by an obstacle, the foragers leave the path to explore new routes. If an ant is successful, it leaves a new trail marking the shortest route on its return. Successful trails are followed by more ants, reinforcing better routes and gradually identifying the best path.[84][85]
Ants use pheromones for more than just making trails. A crushed ant emits an alarm pheromone that sends nearby ants into an attack frenzy and attracts more ants from farther away. Several ant species even use "propaganda pheromones" to confuse enemy ants and make them fight among themselves.[86] Pheromones are produced by a wide range of structures including Dufour's glands, poison glands and glands on the hindgut, pygidium, rectum, sternum, and hind tibia.[67] Pheromones also are exchanged, mixed with food, and passed by trophallaxis, transferring information within the colony.[87] This allows other ants to detect what task group (e.g., foraging or nest maintenance) other colony members belong to.[88] In ant species with queen castes, when the dominant queen stops producing a specific pheromone, workers begin to raise new queens in the colony.[89]
Some ants produce sounds by stridulation, using the gaster segments and their mandibles. Sounds may be used to communicate with colony members or with other species.[90][91]
Defence
See also: Defense in insects
A Plectroctena sp. attacks another of its kind to protect its territory.
Ants attack and defend themselves by biting and, in many species, by stinging often injecting or spraying chemicals. Bullet ants (Paraponera), located in Central and South America, are considered to have the most painful sting of any insect, although it is usually not fatal to humans. This sting is given the highest rating on the Schmidt sting pain index.[92]
The sting of jack jumper ants can be lethal for humans,[93] and an antivenom has been developed for it.[94] Fire ants, Solenopsis spp., are unique in having a venom sac containing piperidine alkaloids.[95] Their stings are painful and can be dangerous to hypersensitive people.[96] Formicine ants secrete a poison from their glands, made mainly of formic acid.[97]
A weaver ant in fighting position, mandibles wide open
Trap-jaw ants of the genus Odontomachus are equipped with mandibles called trap-jaws, which snap shut faster than any other predatory appendages within the animal kingdom.[98] One study of Odontomachus bauri recorded peak speeds of between 126 and 230 km/h (78 and 143 mph), with the jaws closing within 130 microseconds on average. The ants were also observed to use their jaws as a catapult to eject intruders or fling themselves backward to escape a threat.[98] Before striking, the ant opens its mandibles extremely widely and locks them in this position by an internal mechanism. Energy is stored in a thick band of muscle and explosively released when triggered by the stimulation of sensory organs resembling hairs on the inside of the mandibles. The mandibles also permit slow and fine movements for other tasks. Trap-jaws also are seen in other ponerines such as Anochetus, as well as some genera in the tribe Attini, such as Daceton, Orectognathus, and Strumigenys,[98][99] which are viewed as examples of convergent evolution.
A Malaysian species of ant in the Camponotus cylindricus group has enlarged mandibular glands that extend into their gaster. If combat takes a turn for the worse, a worker may perform a final act of suicidal altruism by rupturing the membrane of its gaster, causing the content of its mandibular glands to burst from the anterior region of its head, spraying a poisonous, corrosive secretion containing acetophenones and other chemicals that immobilise small insect attackers. The worker subsequently dies.[100]
Ant mound holes prevent water from entering the nest during rain.
In addition to defence against predators, ants need to protect their colonies from pathogens. Secretions from the metapleural gland, unique to the ants, produce a complex range of chemicals including several with antibiotic properties.[101] Some worker ants maintain the hygiene of the colony and their activities include undertaking or necrophoresis, the disposal of dead nest-mates.[102] Oleic acid has been identified as the compound released from dead ants that triggers necrophoric behaviour in Atta mexicana[103] while workers of Linepithema humile react to the absence of characteristic chemicals (dolichodial and iridomyrmecin) present on the cuticle of their living nestmates to trigger similar behaviour.[104]
Nests may be protected from physical threats such as flooding and overheating by elaborate nest architecture.[105][106] Workers of Cataulacus muticus, an arboreal species that lives in plant hollows, respond to flooding by drinking water inside the nest, and excreting it outside.[107] Camponotus anderseni, which nests in the cavities of wood in mangrove habitats, deals with submergence under water by switching to anaerobic respiration.[108]
Learning
Two Weaver ants walking in tandem
Many animals can learn behaviours by imitation, but ants may be the only group apart from mammals where interactive teaching has been observed. A knowledgeable forager of Temnothorax albipennis can lead a naïve nest-mate to newly discovered food by the process of tandem running. The follower obtains knowledge through its leading tutor. The leader is acutely sensitive to the progress of the follower and slows down when the follower lags and speeds up when the follower gets too close.[109]
Controlled experiments with colonies of Cerapachys biroi suggest that an individual may choose nest roles based on her previous experience. An entire generation of identical workers was divided into two groups whose outcome in food foraging was controlled. One group was continually rewarded with prey, while it was made certain that the other failed. As a result, members of the successful group intensified their foraging attempts while the unsuccessful group ventured out fewer and fewer times. A month later, the successful foragers continued in their role while the others had moved to specialise in brood care.[110]
Nest construction
Main article: Ant colony
Leaf nest of weaver ants, Pamalican, Philippines
Complex nests are built by many ant species, but other species are nomadic and do not build permanent structures. Ants may form subterranean nests or build them on trees. These nests may be found in the ground, under stones or logs, inside logs, hollow stems, or even acorns. The materials used for construction include soil and plant matter,[76] and ants carefully select their nest sites; Temnothorax albipennis will avoid sites with dead ants, as these may indicate the presence of pests or disease. They are quick to abandon established nests at the first sign of threats.[111]
The army ants of South America, such as the Eciton burchellii species, and the driver ants of Africa do not build permanent nests, but instead, alternate between nomadism and stages where the workers form a temporary nest (bivouac) from their own bodies, by holding each other together.[112]
Weaver ant (Oecophylla spp.) workers build nests in trees by attaching leaves together, first pulling them together with bridges of workers and then inducing their larvae to produce silk as they are moved along the leaf edges. Similar forms of nest construction are seen in some species of Polyrhachis.[113]
Ant bridge
Formica polyctena, among other ant species, constructs nests that maintain a relatively constant interior temperature that aids in the development of larvae. The ants maintain the nest temperature by choosing the location, nest materials, controlling ventilation and maintaining the heat from solar radiation, worker activity and metabolism, and in some moist nests, microbial activity in the nest materials.[114][115]
Some ant species, such as those that use natural cavities, can be opportunistic and make use of the controlled micro-climate provided inside human dwellings and other artificial structures to house their colonies and nest structures.[116][117]
Cultivation of food
Main article: Ant–fungus mutualism
Myrmecocystus, honeypot ants, store food to prevent colony famine.
Most ants are generalist predators, scavengers, and indirect herbivores,[16] but a few have evolved specialised ways of obtaining nutrition. It is believed that many ant species that engage in indirect herbivory rely on specialized symbiosis with their gut microbes[118] to upgrade the nutritional value of the food they collect[119] and allow them to survive in nitrogen poor regions, such as rainforest canopies.[120] Leafcutter ants (Atta and Acromyrmex) feed exclusively on a fungus that grows only within their colonies. They continually collect leaves which are taken to the colony, cut into tiny pieces and placed in fungal gardens. Ergates specialise in related tasks according to their sizes. The largest ants cut stalks, smaller workers chew the leaves and the smallest tend the fungus. Leafcutter ants are sensitive enough to recognise the reaction of the fungus to different plant material, apparently detecting chemical signals from the fungus. If a particular type of leaf is found to be toxic to the fungus, the colony will no longer collect it. The ants feed on structures produced by the fungi called gongylidia. Symbiotic bacteria on the exterior surface of the ants produce antibiotics that kill bacteria introduced into the nest that may harm the fungi.[121]
Navigation
An ant trail
Foraging ants travel distances of up to 200 metres (700 ft) from their nest [122] and scent trails allow them to find their way back even in the dark. In hot and arid regions, day-foraging ants face death by desiccation, so the ability to find the shortest route back to the nest reduces that risk. Diurnal desert ants of the genus Cataglyphis such as the Sahara desert ant navigate by keeping track of direction as well as distance travelled. Distances travelled are measured using an internal pedometer that keeps count of the steps taken[123] and also by evaluating the movement of objects in their visual field (optical flow).[124] Directions are measured using the position of the sun.[125]
They integrate this information to find the shortest route back to their nest.[126]
Like all ants, they can also make use of visual landmarks when available[127] as well as olfactory and tactile cues to navigate.[128][129] Some species of ant are able to use the Earth's magnetic field for navigation.[130] The compound eyes of ants have specialised cells that detect polarised light from the Sun, which is used to determine direction.[131][132]
These polarization detectors are sensitive in the ultraviolet region of the light spectrum.[133] In some army ant species, a group of foragers who become separated from the main column may sometimes turn back on themselves and form a circular ant mill. The workers may then run around continuously until they die of exhaustion.[134]
Locomotion
The female worker ants do not have wings and reproductive females lose their wings after their mating flights in order to begin their colonies. Therefore, unlike their wasp ancestors, most ants travel by walking. Some species are capable of leaping. For example, Jerdon's jumping ant (Harpegnathos saltator) is able to jump by synchronising the action of its mid and hind pairs of legs.[135] There are several species of gliding ant including Cephalotes atratus; this may be a common trait among arboreal ants with small colonies. Ants with this ability are able to control their horizontal movement so as to catch tree trunks when they fall from atop the forest canopy.[136]
Other species of ants can form chains to bridge gaps over water, underground, or through spaces in vegetation. Some species also form floating rafts that help them survive floods.[137] These rafts may also have a role in allowing ants to colonise islands.[138] Polyrhachis sokolova, a species of ant found in Australian mangrove swamps, can swim and live in underwater nests. Since they lack gills, they go to trapped pockets of air in the submerged nests to breathe.[139]
Cooperation and competition
Meat-eater ants feeding on a cicada: social ants cooperate and collectively gather food
Not all ants have the same kind of societies. The Australian bulldog ants are among the biggest and most basal of ants. Like virtually all ants, they are eusocial, but their social behaviour is poorly developed compared to other species. Each individual hunts alone, using her large eyes instead of chemical senses to find prey.[140]
Some species attack and take over neighbouring ant colonies. Extreme specialists among these slave-raiding ants, such as the Amazon ants, are incapable of feeding themselves and need captured workers to survive.[141] Captured workers of enslaved Temnothorax species have evolved a counter-strategy, destroying just the female pupae of the slave-making Temnothorax americanus, but sparing the males (who do not take part in slave-raiding as adults).[142]
A worker Harpegnathos saltator (a jumping ant) engaged in battle with a rival colony's queen (on top)
Ants identify kin and nestmates through their scent, which comes from hydrocarbon-laced secretions that coat their exoskeletons. If an ant is separated from its original colony, it will eventually lose the colony scent. Any ant that enters a colony without a matching scent will be attacked.[143]
Parasitic ant species enter the colonies of host ants and establish themselves as social parasites; species such as Strumigenys xenos are entirely parasitic and do not have workers, but instead, rely on the food gathered by their Strumigenys perplexa hosts.[144][145] This form of parasitism is seen across many ant genera, but the parasitic ant is usually a species that is closely related to its host. A variety of methods are employed to enter the nest of the host ant. A parasitic queen may enter the host nest before the first brood has hatched, establishing herself prior to development of a colony scent. Other species use pheromones to confuse the host ants or to trick them into carrying the parasitic queen into the nest. Some simply fight their way into the nest.[146]
A conflict between the sexes of a species is seen in some species of ants with these reproducers apparently competing to produce offspring that are as closely related to them as possible. The most extreme form involves the production of clonal offspring. An extreme of sexual conflict is seen in Wasmannia auropunctata, where the queens produce diploid daughters by thelytokous parthenogenesis and males produce clones by a process whereby a diploid egg loses its maternal contribution to produce haploid males who are clones of the father.[147]
Relationships with other organisms
The spider Myrmarachne plataleoides (female shown) mimics weaver ants to avoid predators.
Ants form symbiotic associations with a range of species, including other ant species, other insects, plants, and fungi. They also are preyed on by many animals and even certain fungi. Some arthropod species spend part of their lives within ant nests, either preying on ants, their larvae, and eggs, consuming the food stores of the ants, or avoiding predators. These inquilines may bear a close resemblance to ants. The nature of this ant mimicry (myrmecomorphy) varies, with some cases involving Batesian mimicry, where the mimic reduces the risk of predation. Others show Wasmannian mimicry, a form of mimicry seen only in inquilines.[148][149]
An ant collects honeydew from an aphid
Aphids and other hemipteran insects secrete a sweet liquid called honeydew, when they feed on plant sap. The sugars in honeydew are a high-energy food source, which many ant species collect.[150] In some cases, the aphids secrete the honeydew in response to ants tapping them with their antennae. The ants in turn keep predators away from the aphids and will move them from one feeding location to another. When migrating to a new area, many colonies will take the aphids with them, to ensure a continued supply of honeydew. Ants also tend mealybugs to harvest their honeydew. Mealybugs may become a serious pest of pineapples if ants are present to protect mealybugs from their natural enemies.[151]
Myrmecophilous (ant-loving) caterpillars of the butterfly family Lycaenidae (e.g., blues, coppers, or hairstreaks) are herded by the ants, led to feeding areas in the daytime, and brought inside the ants' nest at night. The caterpillars have a gland which secretes honeydew when the ants massage them. Some caterpillars produce vibrations and sounds that are perceived by the ants.[152] A similar adaptation can be seen in Grizzled skipper butterflies that emit vibrations by expanding their wings in order to communicate with ants, which are natural predators of these butterflies.[153] Other caterpillars have evolved from ant-loving to ant-eating: these myrmecophagous caterpillars secrete a pheromone that makes the ants act as if the caterpillar is one of their own larvae. The caterpillar is then taken into the ant nest where it feeds on the ant larvae.[154] A number of specialized bacteria have been found as endosymbionts in ant guts. Some of the dominant bacteria belong to the order Hyphomicrobiales whose members are known for being nitrogen-fixing symbionts in legumes but the species found in ant lack the ability to fix nitrogen.[155][156] Fungus-growing ants that make up the tribe Attini, including leafcutter ants, cultivate certain species of fungus in the genera Leucoagaricus or Leucocoprinus of the family Agaricaceae. In this ant-fungus mutualism, both species depend on each other for survival. The ant Allomerus decemarticulatus has evolved a three-way association with the host plant, Hirtella physophora (Chrysobalanaceae), and a sticky fungus which is used to trap their insect prey.[157]
Ants may obtain nectar from flowers such as the dandelion, but are only rarely known to pollinate flowers.
Lemon ants make devil's gardens by killing surrounding plants with their stings and leaving a pure patch of lemon ant trees, (Duroia hirsuta). This modification of the forest provides the ants with more nesting sites inside the stems of the Duroia trees.[158] Although some ants obtain nectar from flowers, pollination by ants is somewhat rare, one example being of the pollination of the orchid Leporella fimbriata which induces male Myrmecia urens to pseudocopulate with the flowers, transferring pollen in the process.[159] One theory that has been proposed for the rarity of pollination is that the secretions of the metapleural gland inactivate and reduce the viability of pollen.[160][161] Some plants have special nectar exuding structures, extrafloral nectaries, that provide food for ants, which in turn protect the plant from more damaging herbivorous insects.[162] Species such as the bullhorn acacia (Acacia cornigera) in Central America have hollow thorns that house colonies of stinging ants (Pseudomyrmex ferruginea) who defend the tree against insects, browsing mammals, and epiphytic vines. Isotopic labelling studies suggest that plants also obtain nitrogen from the ants.[163] In return, the ants obtain food from protein- and lipid-rich Beltian bodies. In Fiji Philidris nagasau (Dolichoderinae) are known to selectively grow species of epiphytic Squamellaria (Rubiaceae) which produce large domatia inside which the ant colonies nest. The ants plant the seeds and the domatia of young seedling are immediately occupied and the ant faeces in them contribute to rapid growth.[164] Similar dispersal associations are found with other dolichoderines in the region as well.[165] Another example of this type of ectosymbiosis comes from the Macaranga tree, which has stems adapted to house colonies of Crematogaster ants.[166]
Many plant species have seeds that are adapted for dispersal by ants.[167] Seed dispersal by ants or myrmecochory is widespread, and new estimates suggest that nearly 9% of all plant species may have such ant associations.[168][167] Often, seed-dispersing ants perform directed dispersal, depositing the seeds in locations that increase the likelihood of seed survival to reproduction.[169] Some plants in arid, fire-prone systems are particularly dependent on ants for their survival and dispersal as the seeds are transported to safety below the ground.[170] Many ant-dispersed seeds have special external structures, elaiosomes, that are sought after by ants as food.[171] Ants can substantially alter rate of decomposition and nutrient cycling in their nest.[172][173] By myrmecochory and modification of soil conditions they substantially alter vegetation and nutrient cycling in surrounding ecosystem.[174]
A convergence, possibly a form of mimicry, is seen in the eggs of stick insects. They have an edible elaiosome-like structure and are taken into the ant nest where the young hatch.[175]
A meat ant tending a common leafhopper nymph
Bold Jumping Spider (Phidippus audax) with a cutworm (tribe Noctuini) and then lost to ants (Family Formicidae)
ants from different colonies steal the cranefly that a pair of Long-jawed orb weaver spiders were consuming.
Most ants are predatory and some prey on and obtain food from other social insects including other ants. Some species specialise in preying on termites (Megaponera and Termitopone) while a few Cerapachyinae prey on other ants.[122] Some termites, including Nasutitermes corniger, form associations with certain ant species to keep away predatory ant species.[176] The tropical wasp Mischocyttarus drewseni coats the pedicel of its nest with an ant-repellent chemical.[177] It is suggested that many tropical wasps may build their nests in trees and cover them to protect themselves from ants. Other wasps, such as A. multipicta, defend against ants by blasting them off the nest with bursts of wing buzzing.[178] Stingless bees (Trigona and Melipona) use chemical defences against ants.[122]
Flies in the Old World genus Bengalia (Calliphoridae) prey on ants and are kleptoparasites, snatching prey or brood from the mandibles of adult ants.[179] Wingless and legless females of the Malaysian phorid fly (Vestigipoda myrmolarvoidea) live in the nests of ants of the genus Aenictus and are cared for by the ants.[179]
Oecophylla smaragdina killed by a fungus
Fungi in the genera Cordyceps and Ophiocordyceps infect ants. Ants react to their infection by climbing up plants and sinking their mandibles into plant tissue. The fungus kills the ants, grows on their remains, and produces a fruiting body. It appears that the fungus alters the behaviour of the ant to help disperse its spores [180] in a microhabitat that best suits the fungus.[181] Strepsipteran parasites also manipulate their ant host to climb grass stems, to help the parasite find mates.[182]
A nematode (Myrmeconema neotropicum) that infects canopy ants (Cephalotes atratus) causes the black-coloured gasters of workers to turn red. The parasite also alters the behaviour of the ant, causing them to carry their gasters high. The conspicuous red gasters are mistaken by birds for ripe fruits, such as Hyeronima alchorneoides, and eaten. The droppings of the bird are collected by other ants and fed to their young, leading to further spread of the nematode.[183]
Spiders (Like this Menemerus jumping spider) sometimes feed on ants
A study of Temnothorax nylanderi colonies in Germany found that workers parasitized by the tapeworm Anomotaenia brevis (ants are intermediate hosts, the definitive hosts are woodpeckers) lived much longer than unparasitized workers and had a reduced mortality rate, comparable to that of the queens of the same species, which live for as long as two decades.[184]
South American poison dart frogs in the genus Dendrobates feed mainly on ants, and the toxins in their skin may come from the ants.[185]
Army ants forage in a wide roving column, attacking any animals in that path that are unable to escape. In Central and South America, Eciton burchellii is the swarming ant most commonly attended by "ant-following" birds such as antbirds and woodcreepers.[186][187] This behaviour was once considered mutualistic, but later studies found the birds to be parasitic. Direct kleptoparasitism (birds stealing food from the ants' grasp) is rare and has been noted in Inca doves which pick seeds at nest entrances as they are being transported by species of Pogonomyrmex.[188] Birds that follow ants eat many prey insects and thus decrease the foraging success of ants.[189] Birds indulge in a peculiar behaviour called anting that, as yet, is not fully understood. Here birds rest on ant nests, or pick and drop ants onto their wings and feathers; this may be a means to remove ectoparasites from the birds.
Anteaters, aardvarks, pangolins, echidnas and numbats have special adaptations for living on a diet of ants. These adaptations include long, sticky tongues to capture ants and strong claws to break into ant nests. Brown bears (Ursus arctos) have been found to feed on ants. About 12%, 16%, and 4% of their faecal volume in spring, summer and autumn, respectively, is composed of ants.[190]
Relationship with humans
Weaver ants are used as a biological control for citrus cultivation in southern China.
Ants perform many ecological roles that are beneficial to humans, including the suppression of pest populations and aeration of the soil. The use of weaver ants in citrus cultivation in southern China is considered one of the oldest known applications of biological control.[191] On the other hand, ants may become nuisances when they invade buildings or cause economic losses.
In some parts of the world (mainly Africa and South America), large ants, especially army ants, are used as surgical sutures. The wound is pressed together and ants are applied along it. The ant seizes the edges of the wound in its mandibles and locks in place. The body is then cut off and the head and mandibles remain in place to close the wound.[192][193][194] The large heads of the dinergates (soldiers) of the leafcutting ant Atta cephalotes are also used by native surgeons in closing wounds.[195]
Some ants have toxic venom and are of medical importance. The species include Paraponera clavata (tocandira) and Dinoponera spp. (false tocandiras) of South America [196] and the Myrmecia ants of Australia.[197]
In South Africa, ants are used to help harvest the seeds of rooibos (Aspalathus linearis), a plant used to make a herbal tea. The plant disperses its seeds widely, making manual collection difficult. Black ants collect and store these and other seeds in their nest, where humans can gather them en masse. Up to half a pound (200 g) of seeds may be collected from one ant-heap.[198][199]
Although most ants survive attempts by humans to eradicate them, a few are highly endangered. These tend to be island species that have evolved specialized traits and risk being displaced by introduced ant species. Examples include the critically endangered Sri Lankan relict ant (Aneuretus simoni) and Adetomyrma venatrix of Madagascar.[200]
As food
See also: Entomophagy
Roasted ants in Colombia
Ant larvae for sale in Isaan, Thailand
Ants and their larvae are eaten in different parts of the world. The eggs of two species of ants are used in Mexican escamoles. They are considered a form of insect caviar and can sell for as much as US$50 per kg going up to US$200 per kg (as of 2006) because they are seasonal and hard to find.[201] In the Colombian department of Santander, hormigas culonas (roughly interpreted as "large-bottomed ants") Atta laevigata are toasted alive and eaten.[202] In areas of India, and throughout Burma and Thailand, a paste of the green weaver ant (Oecophylla smaragdina) is served as a condiment with curry.[203] Weaver ant eggs and larvae, as well as the ants, may be used in a Thai salad, yam (Thai: ยำ), in a dish called yam khai mot daeng (Thai: ยำไข่มดแดง) or red ant egg salad, a dish that comes from the Issan or north-eastern region of Thailand. Saville-Kent, in the Naturalist in Australia wrote "Beauty, in the case of the green ant, is more than skin-deep. Their attractive, almost sweetmeat-like translucency possibly invited the first essays at their consumption by the human species". Mashed up in water, after the manner of lemon squash, "these ants form a pleasant acid drink which is held in high favor by the natives of North Queensland, and is even appreciated by many European palates".[204]
In his First Summer in the Sierra, John Muir notes that the Digger Indians of California ate the tickling, acid gasters of the large jet-black carpenter ants. The Mexican Indians eat the repletes, or living honey-pots, of the honey ant (Myrmecocystus).[204]
As pests
See also: Ants of medical importance
The tiny pharaoh ant is a major pest in hospitals and office blocks; it can make nests between sheets of paper.
Some ant species are considered as pests, primarily those that occur in human habitations, where their presence is often problematic. For example, the presence of ants would be undesirable in sterile places such as hospitals or kitchens. Some species or genera commonly categorized as pests include the Argentine ant, immigrant pavement ant, yellow crazy ant, banded sugar ant, pharaoh ant, red wood ant, black carpenter ant, odorous house ant, red imported fire ant, and European fire ant. Some ants will raid stored food, some will seek water sources, others may damage indoor structures, some may damage agricultural crops directly or by aiding sucking pests. Some will sting or bite.[205] The adaptive nature of ant colonies make it nearly impossible to eliminate entire colonies and most pest management practices aim to control local populations and tend to be temporary solutions. Ant populations are managed by a combination of approaches that make use of chemical, biological, and physical methods. Chemical methods include the use of insecticidal bait which is gathered by ants as food and brought back to the nest where the poison is inadvertently spread to other colony members through trophallaxis. Management is based on the species and techniques may vary according to the location and circumstance.[205]
In science and technology
See also: Myrmecology, Biomimetics, and Ant colony optimization algorithms
Camponotus nearcticus workers travelling between two formicaria through connector tubing
Observed by humans since the dawn of history, the behaviour of ants has been documented and the subject of early writings and fables passed from one century to another. Those using scientific methods, myrmecologists, study ants in the laboratory and in their natural conditions. Their complex and variable social structures have made ants ideal model organisms. Ultraviolet vision was first discovered in ants by Sir John Lubbock in 1881.[206] Studies on ants have tested hypotheses in ecology and sociobiology, and have been particularly important in examining the predictions of theories of kin selection and evolutionarily stable strategies.[207] Ant colonies may be studied by rearing or temporarily maintaining them in formicaria, specially constructed glass framed enclosures.[208] Individuals may be tracked for study by marking them with dots of colours.[209]
The successful techniques used by ant colonies have been studied in computer science and robotics to produce distributed and fault-tolerant systems for solving problems, for example Ant colony optimization and Ant robotics. This area of biomimetics has led to studies of ant locomotion, search engines that make use of "foraging trails", fault-tolerant storage, and networking algorithms.[210]
As pets
Main article: Ant-keeping
From the late 1950s through the late 1970s, ant farms were popular educational children's toys in the United States. Some later commercial versions use transparent gel instead of soil, allowing greater visibility at the cost of stressing the ants with unnatural light.[211]
In culture
Aesop's ants: illustration by Milo Winter, 1888–1956
Anthropomorphised ants have often been used in fables and children's stories to represent industriousness and cooperative effort. They also are mentioned in religious texts.[212][213] In the Book of Proverbs in the Bible, ants are held up as a good example of hard work and cooperation.[214] Aesop did the same in his fable The Ant and the Grasshopper. In the Quran, Sulayman is said to have heard and understood an ant warning other ants to return home to avoid being accidentally crushed by Sulayman and his marching army.[Quran 27:18],[215][216] In parts of Africa, ants are considered to be the messengers of the deities. Some Native American mythology, such as the Hopi mythology, considers ants as the very first animals. Ant bites are often said to have curative properties. The sting of some species of Pseudomyrmex is claimed to give fever relief.[217] Ant bites are used in the initiation ceremonies of some Amazon Indian cultures as a test of endurance.[218][219] In Greek mythology, the goddess Athena turned the maiden Myrmex into an ant when the latter claimed to have invented the plough, when in fact it was Athena's own invention.[220]
An ant pictured in the coat of arms of Multia, a town in Finland
Ant society has always fascinated humans and has been written about both humorously and seriously. Mark Twain wrote about ants in his 1880 book A Tramp Abroad.[221] Some modern authors have used the example of the ants to comment on the relationship between society and the individual. Examples are Robert Frost in his poem "Departmental" and T. H. White in his fantasy novel The Once and Future King. The plot in French entomologist and writer Bernard Werber's Les Fourmis science-fiction trilogy is divided between the worlds of ants and humans; ants and their behaviour are described using contemporary scientific knowledge. H.G. Wells wrote about intelligent ants destroying human settlements in Brazil and threatening human civilization in his 1905 science-fiction short story, The Empire of the Ants. A similar German story involving army ants, Leiningen Versus the Ants, was written in 1937 and recreated in movie form as The Naked Jungle in 1954.[222] In more recent times, animated cartoons and 3-D animated films featuring ants have been produced including Antz, A Bug's Life, The Ant Bully, The Ant and the Aardvark, Ferdy the Ant and Atom Ant. Renowned myrmecologist E. O. Wilson wrote a short story, "Trailhead" in 2010 for The New Yorker magazine, which describes the life and death of an ant-queen and the rise and fall of her colony, from an ants' point of view.[223] The French neuroanatomist, psychiatrist and eugenicist Auguste Forel believed that ant societies were models for human society. He published a five volume work from 1921 to 1923 that examined ant biology and society.[224]
In the early 1990s, the video game SimAnt, which simulated an ant colony, won the 1992 Codie award for "Best Simulation Program".[225]
Ants also are quite popular inspiration for many science-fiction insectoids, such as the Formics of Ender's Game, the Bugs of Starship Troopers, the giant ants in the films Them! and Empire of the Ants, Marvel Comics' super hero Ant-Man, and ants mutated into super-intelligence in Phase IV. In computer strategy games, ant-based species often benefit from increased production rates due to their single-minded focus, such as the Klackons in the Master of Orion series of games or the ChCht in Deadlock II. These characters are often credited with a hive mind, a common misconception about ant colonies.[226]
See also
Main article: Outline of ants
Glossary of ant terms
International Union for the Study of Social Insects
Myrmecological News (journal)
Task allocation and partitioning in social insects
References
^ a b Moreau CS, Bell CD, Vila R, Archibald SB, Pierce NE (April 2006). "Phylogeny of the ants: diversification in the age of angiosperms". Science. 312 (5770): 101–104. Bibcode:2006Sci...312..101M. doi:10.1126/science.1124891. PMID 16601190. S2CID 20729380.
^ Borowiec, Marek L.; Moreau, Corrie S.; Rabeling, Christian (2020). "Ants: Phylogeny and Classification". In Starr, Christopher K. (ed.). Encyclopedia of Social Insects. Cham: Springer International Publishing. pp. 1–18. doi:10.1007/978-3-319-90306-4_155-1. ISBN 978-3-319-90306-4. S2CID 219873464. Retrieved 11 February 2024.
^ a b Ward PS (2007). "Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae)" (PDF). Zootaxa. 1668: 549–563. doi:10.11646/zootaxa.1668.1.26. Archived (PDF) from the original on 9 October 2022.
^ a b Rabeling C, Brown JM, Verhaagh M (September 2008). "Newly discovered sister lineage sheds light on early ant evolution". Proceedings of the National Academy of Sciences of the United States of America. 105 (39): 14913–14917. Bibcode:2008PNAS..10514913R. doi:10.1073/pnas.0806187105. PMC 2567467. PMID 18794530.
^ Brady SG, Fisher BL, Schultz TR, Ward PS (May 2014). "The rise of army ants and their relatives: diversification of specialized predatory doryline ants". BMC Evolutionary Biology. 14 (93): 93. Bibcode:2014BMCEE..14...93B. doi:10.1186/1471-2148-14-93. PMC 4021219. PMID 24886136.
^ emmet. Merriam-Webster Dictionary
^ "ant". Merriam-Webster Online Dictionary. Retrieved 6 June 2008.
^ "Ant. Online Etymology Dictionary". Retrieved 30 May 2009.
^ Simpson DP (1979). Cassell's Latin Dictionary (5th ed.). London: Cassell. ISBN 978-0-304-52257-6.
^ "Formic". Etymonline.com. Retrieved 30 January 2012.
^ "Pismire". Etymonline.com. Retrieved 27 August 2020.
^ a b Johnson BR, Borowiec ML, Chiu JC, Lee EK, Atallah J, Ward PS (October 2013). "Phylogenomics resolves evolutionary relationships among ants, bees, and wasps". Current Biology. 23 (20): 2058–2062. Bibcode:2013CBio...23.2058J. doi:10.1016/j.cub.2013.08.050. PMID 24094856.
^ Wilson EO, Carpenter FM, Brown WL (September 1967). "The first mesozoic ants". Science. 157 (3792): 1038–1040. Bibcode:1967Sci...157.1038W. doi:10.1126/science.157.3792.1038. PMID 17770424. S2CID 43155424.
^ Boudinot, Brendon E; Richter, Adrian; Katzke, Julian; Chaul, Júlio C M; Keller, Roberto A; Economo, Evan P; Beutel, Rolf Georg; Yamamoto, Shûhei (29 July 2022). "Evidence for the evolution of eusociality in stem ants and a systematic revision of † Gerontoformica (Hymenoptera: Formicidae)". Zoological Journal of the Linnean Society. 195 (4): 1355–1389. doi:10.1093/zoolinnean/zlab097. hdl:10451/55807. ISSN 0024-4082.
^ a b Grimaldi D, Agosti D (December 2000). "A formicine in New Jersey cretaceous amber (Hymenoptera: formicidae) and early evolution of the ants". Proceedings of the National Academy of Sciences of the United States of America. 97 (25): 13678–13683. Bibcode:2000PNAS...9713678G. doi:10.1073/pnas.240452097. PMC 17635. PMID 11078527.
^ a b Wilson EO, Hölldobler B (May 2005). "The rise of the ants: a phylogenetic and ecological explanation". Proceedings of the National Academy of Sciences of the United States of America. 102 (21): 7411–7414. Bibcode:2005PNAS..102.7411W. doi:10.1073/pnas.0502264102. PMC 1140440. PMID 15899976.
^ LaPolla JS, Dlussky GM, Perrichot V (2013). "Ants and the fossil record". Annual Review of Entomology. 58: 609–730. doi:10.1146/annurev-ento-120710-100600. PMID 23317048. S2CID 40555356.
^ Barden P, Grimaldi D (2012). "Rediscovery of the bizarre Cretaceous ant Haidomyrmex Dlussky (Hymenoptera: Formicidae), with two new species" (PDF). American Museum Novitates (3755): 1–16. doi:10.1206/3755.2. hdl:2246/6368. S2CID 83598305. Archived from the original (PDF) on 23 April 2013. Retrieved 5 May 2013.
^ Hölldobler & Wilson (1990), pp. 23–24
^ Thorne BL (1997). "Evolution of eusociality in termites" (PDF). Annu. Rev. Ecol. Syst. 28 (5): 27–53. doi:10.1146/annurev.ecolsys.28.1.27. PMC 349550. Archived from the original (PDF) on 30 May 2010.
^ "Order Isoptera – Termites". Iowa State University Entomology. 16 February 2004. Archived from the original on 15 June 2008. Retrieved 12 June 2008.
^ "Family Mutillidae – Velvet ants". Iowa State University Entomology. 16 February 2004. Archived from the original on 30 June 2008. Retrieved 12 June 2008.
^ Hölldobler & Wilson (1990), p. 4
^ Jones, Alice S. "Fantastic ants – Did you know?". National Geographic Magazine. Archived from the original on 30 July 2008. Retrieved 5 July 2008.
^ Thomas, Philip (2007). "Pest Ants in Hawaii". Hawaiian Ecosystems at Risk project (HEAR). Retrieved 6 July 2008.
^ Fayle, Tom M.; Klimes, Petr (18 October 2022). "Improving estimates of global ant biomass and abundance". Proceedings of the National Academy of Sciences. 119 (42): e2214825119. Bibcode:2022PNAS..11914825F. doi:10.1073/pnas.2214825119. ISSN 0027-8424. PMC 9586285. PMID 36197959.
^ Holldobler B, Wilson EO (2009). The Superorganism: The Beauty, Elegance, and Strangeness of Insect Societies. New York: W.W. Norton. p. 5. ISBN 978-0-393-06704-0.
^ Schultz TR (December 2000). "In search of ant ancestors". Proceedings of the National Academy of Sciences of the United States of America. 97 (26): 14028–14029. Bibcode:2000PNAS...9714028S. doi:10.1073/pnas.011513798. PMC 34089. PMID 11106367.
^ "How many ants are there for every one person on earth?". info.com. Archived from the original on 13 August 2013. Retrieved 27 July 2013.
^ Schultheiss, Patrick; Nooten, Sabine S.; Wang, Runxi; Wong, Mark K. L.; Brassard, François; Guénard, Benoit (4 October 2022). "The abundance, biomass, and distribution of ants on Earth". Proceedings of the National Academy of Sciences. 119 (40): e2201550119. Bibcode:2022PNAS..11901550S. doi:10.1073/pnas.2201550119. ISSN 0027-8424. PMC 9546634. PMID 36122199. S2CID 252381912.
^ Hölldobler & Wilson (1990), p. 589
^ Shattuck SO (1999). Australian ants: their biology and identification. Collingwood, Vic: CSIRO. p. 149. ISBN 978-0-643-06659-5.
^ Schaal, Stephan (27 January 2006). "Messel". Encyclopedia of Life Sciences. doi:10.1038/npg.els.0004143. ISBN 978-0-470-01617-6.
^ a b AntWeb
^ Agosti D, Majer JD, Alonso JE, Schultz TR, eds. (2000). Ants: Standard methods for measuring and monitoring biodiversity. Smithsonian Institution Press. Retrieved 13 December 2015.
^ Johnson NF (2007). "Hymenoptera name server". Ohio State University. Archived from the original on 27 January 2016. Retrieved 6 July 2008.
^ Borror, Triplehorn & Delong (1989), p. 737
^ Ouellette, Gary D.; Fisher, Brian L.; Girman, Derek J. (2006). "Molecular systematics of basal subfamilies of ants using 28S rRNA (Hymenoptera: Formicidae)". Molecular Phylogenetics and Evolution. 40 (2): 359–369. doi:10.1016/j.ympev.2006.03.017. hdl:10211.1/1549. ISSN 1055-7903. PMID 16630727.
^ a b c d Borror, Triplehorn & Delong (1989), pp. 24–71
^ Fent K, Wehner R (April 1985). "Oceili: a celestial compass in the desert ant cataglyphis". Science. 228 (4696): 192–194. Bibcode:1985Sci...228..192F. doi:10.1126/science.228.4696.192. PMID 17779641. S2CID 33242108.
^ Palavalli-Nettimi R, Narendra A (April 2018). "Miniaturisation decreases visual navigational competence in ants". The Journal of Experimental Biology. 221 (Pt 7): jeb177238. doi:10.1242/jeb.177238. PMID 29487158.
^ Eriksson ES (1985). "Attack behaviour and distance perception in the Australian bulldog ant Myrmecia nigriceps" (PDF). Journal of Experimental Biology. 119 (1): 115–131. doi:10.1242/jeb.119.1.115. Archived (PDF) from the original on 9 October 2022.
^ Eisner T, Happ GM (1962). "The infrabuccal pocket of a formicine ant: a social filtration device". Psyche: A Journal of Entomology. 69 (3): 107–116. doi:10.1155/1962/25068.
^ Holbrook, Tate (22 September 2009). "Ask a Biologist: Face to Face with Ants". ASU School of Life Sciences. Retrieved 23 January 2018.
^ Singh R (2006). Elements of Entomology. Rastogi Publications. p. 284. ISBN 978-8171336777.
^ Wilson EO (June 1953). "The origin and evolution of polymorphism in ants". The Quarterly Review of Biology. 28 (2): 136–156. doi:10.1086/399512. PMID 13074471. S2CID 4560071.
^ Weber, NA (1946). "Dimorphism in the African Oecophylla worker and an anomaly (Hym.: Formicidae)" (PDF). Annals of the Entomological Society of America. 39: 7–10. doi:10.1093/aesa/39.1.7. Archived (PDF) from the original on 9 October 2022.
^ Wilson EO, Taylor RW (1964). "A Fossil Ant Colony: New Evidence of Social Antiquity" (PDF). Psyche: A Journal of Entomology. 71 (2): 93–103. doi:10.1155/1964/17612. Archived (PDF) from the original on 9 October 2022.
^ Moffett MW, Tobin JE (1991). "Physical castes in ant workers: a problem for Daceton armigerum and other ants" (PDF). Psyche: A Journal of Entomology. 98 (4): 283–292. doi:10.1155/1991/30265. Archived from the original (PDF) on 27 February 2008.
^ Børgesen LW (2000). "Nutritional function of replete workers in the pharaoh's ant, Monomorium pharaonis (L.)". Insectes Sociaux. 47 (2): 141–146. doi:10.1007/PL00001692. S2CID 31953751.
^ Rissing SW (1984). "Replete caste production and allometry of workers in the Honey Ant, Myrmecocystus mexicanus Wesmael (Hymenoptera: Formicidae)". Journal of the Kansas Entomological Society. 57 (2): 347–350.
^ Hughes WO, Sumner S, Van Borm S, Boomsma JJ (August 2003). "Worker caste polymorphism has a genetic basis in Acromyrmex leaf-cutting ants". Proceedings of the National Academy of Sciences of the United States of America. 100 (16): 9394–9397. Bibcode:2003PNAS..100.9394H. doi:10.1073/pnas.1633701100. PMC 170929. PMID 12878720.
^ Ross KG, Krieger MJ, Shoemaker DD (December 2003). "Alternative genetic foundations for a key social polymorphism in fire ants". Genetics. 165 (4): 1853–1867. doi:10.1093/genetics/165.4.1853. PMC 1462884. PMID 14704171.
^ Crosland MW, Crozier RH (March 1986). "Myrmecia pilosula, an ant with only one Pair of chromosomes". Science. 231 (4743): 1278. Bibcode:1986Sci...231.1278C. doi:10.1126/science.231.4743.1278. PMID 17839565. S2CID 25465053.
^ Tsutsui ND, Suarez AV, Spagna JC, Johnston JS (February 2008). "The evolution of genome size in ants". BMC Evolutionary Biology. 8 (64): 64. Bibcode:2008BMCEE...8...64T. doi:10.1186/1471-2148-8-64. PMC 2268675. PMID 18302783.
^ a b Moura, Mariana Neves; Cardoso, Danon Clemes; Cristiano, Maykon Passos (2021). "The tight genome size of ants: diversity and evolution under ancestral state reconstruction and base composition". Zoological Journal of the Linnean Society. 193 (1): 124–144. doi:10.1093/zoolinnean/zlaa135. ISSN 0024-4082.
^ Hölldobler B, Wilson EO (1990). The Ants. Harvard University Press. p. 291. ISBN 978-0-674-04075-5.
^ Gillott, Cedric (1995). Entomology. Springer. p. 325. ISBN 978-0-306-44967-3.
^ Anderson KE, Linksvayer TA, Smith CR (2008). "The causes and consequences of genetic caste determination in ants (Hymenoptera: Formicidae)". Myrmecol. News. 11: 119–132.
^ Gaul AT (1951). "A Glossary of Terms and Phrases Used in the Study of Social Insects". Annals of the Entomological Society of America. 44 (3): 473–484. doi:10.1093/aesa/44.3.473. ISSN 1938-2901.
^ Hölldobler & Wilson (1990), pp. 351, 372
^ Traniello JFA (1989). "Foraging strategies of ants". Annual Review of Entomology. 34: 191–210. doi:10.1146/annurev.en.34.010189.001203.
^ Sorensen A, Busch TM, Vinson SB (1984). "Behavioral flexibility of temporal sub-castes in the fire ant, Solenopsis invicta, in response to food". Psyche: A Journal of Entomology. 91 (3–4): 319–332. doi:10.1155/1984/39236.
^ Tofilski, Adam; Couvillon, Margaret J.; Evison, Sophie E. F.; Helanterä, Heikki; Robinson, Elva J. H.; Ratnieks, Francis L. W. (2008). "Preemptive Defensive Self-Sacrifice by Ant Workers". The American Naturalist. 172 (5): E239–E243. doi:10.1086/591688. ISSN 0003-0147. PMID 18928332. S2CID 7052340.
^ Shorter, J. R.; Rueppell, O. (2012). "A review on self-destructive defense behaviors in social insects". Insectes Sociaux. 59 (1): 1–10. doi:10.1007/s00040-011-0210-x. ISSN 0020-1812. S2CID 253634662.
^ Keller L (1998). "Queen lifespan and colony characteristics in ants and termites". Insectes Sociaux. 45 (3): 235–246. doi:10.1007/s000400050084. S2CID 24541087.
^ a b Franks NR, Resh VH, Cardé RT, eds. (2003). Encyclopedia of Insects. San Diego: Academic Press. pp. 29–32. ISBN 978-0-12-586990-4.
^ Kipyatkov VE (2001). "Seasonal life cycles and the forms of dormancy in ants (Hymenoptera, Formicoidea)". Acta Societatis Zoologicae Bohemicae. 65 (2): 198–217.
^ Heinze J, Tsuji K (1995). "Ant reproductive strategies" (PDF). Res. Popul. Ecol. 37 (2): 135–149. Bibcode:1995PopEc..37..135H. doi:10.1007/BF02515814. S2CID 21948488. Archived from the original (PDF) on 27 May 2011. Retrieved 16 April 2009.
^ Mikheyev AS (2003). "Evidence for mating plugs in the fire ant Solenopsis invicta". Insectes Sociaux. 50 (4): 401–402. doi:10.1007/s00040-003-0697-x. S2CID 43492133.
^ Peeters C, Hölldobler B (November 1995). "Reproductive cooperation between queens and their mated workers: the complex life history of an ant with a valuable nest". Proceedings of the National Academy of Sciences of the United States of America. 92 (24): 10977–10979. Bibcode:1995PNAS...9210977P. doi:10.1073/pnas.92.24.10977. PMC 40553. PMID 11607589.
^ Franks NR, Hölldobler B (1987). "Sexual competition during colony reproduction in army ants". Biological Journal of the Linnean Society. 30 (3): 229–243. doi:10.1111/j.1095-8312.1987.tb00298.x.
^ Yamauchi K, Kawase N (1992). "Pheromonal manipulation of workers by a fighting male to kill his rival males in the ant Cardiocondyla wroughtonii". Naturwissenschaften. 79 (6): 274–276. Bibcode:1992NW.....79..274Y. doi:10.1007/BF01175395. S2CID 31191187.
^ Taylor RW (2007). "Bloody funny wasps! Speculations on the evolution of eusociality in ants" (PDF). In Snelling RR, Fisher BL, Ward PS (eds.). Advances in ant systematics (Hymenoptera: Formicidae): homage to E. O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute, 80. American Entomological Institute. pp. 580–609. Retrieved 13 December 2015.
^ Wilson EO (1957). "The organization of a nuptial flight of the ant Pheidole sitarches Wheeler" (PDF). Psyche: A Journal of Entomology. 64 (2): 46–50. doi:10.1155/1957/68319.
^ a b c Hölldobler & Wilson (1990), pp. 143–179
^ Vidal M, Königseder F, Giehr J, Schrempf A, Lucas C, Heinze J (May 2021). "Worker ants promote outbreeding by transporting young queens to alien nests". Communications Biology. 4 (1): 515. doi:10.1038/s42003-021-02016-1. PMC 8093424. PMID 33941829.
^ Sudd JH, Franks NR (2013). The Behavioural Ecology of Ants. Springer Science & Business Media. p. 41. ISBN 978-9400931237.
^ Cook, Zoe; Franks, Daniel W.; Robinson, Elva J.H. (2013). "Exploration versus exploitation in polydomous ant colonies". Journal of Theoretical Biology. 323: 49–56. Bibcode:2013JThBi.323...49C. doi:10.1016/j.jtbi.2013.01.022. PMID 23380232.
^ Bos, Nick; d’Ettorre, Patrizia (2012). "Recognition of Social Identity in Ants". Frontiers in Psychology. 3: 83. doi:10.3389/fpsyg.2012.00083. ISSN 1664-1078. PMC 3309994. PMID 22461777.
^ Moffett, Mark W. (2012). "Supercolonies of billions in an invasive ant: What is a society?". Behavioral Ecology. 23 (5): 925–933. doi:10.1093/beheco/ars043. ISSN 1465-7279.
^ Van Wilgenburg, Ellen; Torres, Candice W.; Tsutsui, Neil D. (2010). "The global expansion of a single ant supercolony". Evolutionary Applications. 3 (2): 136–143. doi:10.1111/j.1752-4571.2009.00114.x. ISSN 1752-4571. PMC 3352483. PMID 25567914.
^ Helanterä, Heikki; Strassmann, Joan E.; Carrillo, Juli; Queller, David C. (2009). "Unicolonial ants: where do they come from, what are they and where are they going?". Trends in Ecology & Evolution. 24 (6): 341–349. doi:10.1016/j.tree.2009.01.013. PMID 19328589.
^ a b Jackson DE, Ratnieks FL (August 2006). "Communication in ants". Current Biology. 16 (15): R570–R574. Bibcode:2006CBio...16.R570J. doi:10.1016/j.cub.2006.07.015. PMID 16890508. S2CID 5835320.
^ Goss S, Aron S, Deneubourg JL, Pasteels JM (1989). "Self-organized shortcuts in the Argentine ant" (PDF). Naturwissenschaften. 76 (12): 579–581. Bibcode:1989NW.....76..579G. doi:10.1007/BF00462870. S2CID 18506807. Archived (PDF) from the original on 9 October 2022.
^ D'Ettorre P, Heinze J (2001). "Sociobiology of slave-making ants". Acta Ethologica. 3 (2): 67–82. doi:10.1007/s102110100038. S2CID 37840769.
^ Detrain C, Deneubourg JL, Pasteels JM (1999). Information processing in social insects. Birkhäuser. pp. 224–227. ISBN 978-3-7643-5792-4.
^ Greene MJ, Gordon DM (March 2007). "Structural complexity of chemical recognition cues affects the perception of group membership in the ants Linephithema humile and Aphaenogaster cockerelli". The Journal of Experimental Biology. 210 (Pt 5): 897–905. doi:10.1242/jeb.02706. PMID 17297148. S2CID 14909476.
^ Hölldobler & Wilson (1990), p. 354
^ Hickling R, Brown RL (October 2000). "Analysis of acoustic communication by ants". The Journal of the Acoustical Society of America. 108 (4): 1920–1929. Bibcode:2000ASAJ..108.1920H. doi:10.1121/1.1290515. PMID 11051518.
^ Roces F, Hölldobler B (1996). "Use of stridulation in foraging leaf-cutting ants: Mechanical support during cutting or short-range recruitment signal?". Behavioral Ecology and Sociobiology. 39 (5): 293–299. doi:10.1007/s002650050292. S2CID 32884747.
^ Schmidt JO, Blum MS, Overal WL (1983). "Hemolytic activities of stinging insect venoms". Archives of Insect Biochemistry and Physiology. 1 (2): 155–160. doi:10.1002/arch.940010205.
^ Clarke PS (1986). "The natural history of sensitivity to jack jumper ants (Hymenoptera formicidae Myrmecia pilosula) in Tasmania". The Medical Journal of Australia. 145 (11–12): 564–566. doi:10.5694/j.1326-5377.1986.tb139498.x. PMID 3796365.
^ Brown SG, Heddle RJ, Wiese MD, Blackman KE (August 2005). "Efficacy of ant venom immunotherapy and whole body extracts". The Journal of Allergy and Clinical Immunology. 116 (2): 464–465, author reply 465–466. doi:10.1016/j.jaci.2005.04.025. PMID 16083810.
^ Obin MS, Vander Meer RK (December 1985). "Gaster flagging by fire ants (Solenopsis spp.): Functional significance of venom dispersal behavior". Journal of Chemical Ecology. 11 (12): 1757–1768. Bibcode:1985JCEco..11.1757O. doi:10.1007/BF01012125. PMID 24311339. S2CID 12182722.
^ Stafford CT (August 1996). "Hypersensitivity to fire ant venom". Annals of Allergy, Asthma & Immunology. 77 (2): 87–95, quiz 96–99. doi:10.1016/S1081-1206(10)63493-X. PMID 8760773.
^ Lopez LC, Morgan ED, Brand JM (1993). "Hexadecanol and hexadecyl formate in the venom gland of formicine ants". Philosophical Transactions of the Royal Society B. 341 (1296): 177–180. doi:10.1098/rstb.1993.0101. S2CID 85361145.
^ a b c Patek SN, Baio JE, Fisher BL, Suarez AV (August 2006). "Multifunctionality and mechanical origins: ballistic jaw propulsion in trap-jaw ants". Proceedings of the National Academy of Sciences of the United States of America. 103 (34): 12787–12792. Bibcode:2006PNAS..10312787P. doi:10.1073/pnas.0604290103. PMC 1568925. PMID 16924120.
^ Gronenberg W (1996). "The trap-jaw mechanism in the dacetine ants Daceton armigerum and Strumigenys sp" (PDF). The Journal of Experimental Biology. 199 (Pt 9): 2021–2033. doi:10.1242/jeb.199.9.2021. PMID 9319931. Archived (PDF) from the original on 9 October 2022.
^ Jones TH, Clark DA, Edwards AA, Davidson DW, Spande TF, Snelling RR (August 2004). "The chemistry of exploding ants, Camponotus spp. (cylindricus complex)". Journal of Chemical Ecology. 30 (8): 1479–1492. Bibcode:2004JCEco..30.1479J. doi:10.1023/B:JOEC.0000042063.01424.28. PMID 15537154. S2CID 23756265.
^ Yek, Sze Huei; Mueller, Ulrich G. (2011). "The metapleural gland of ants" (PDF). Biological Reviews. 86 (4): 774–791. doi:10.1111/j.1469-185X.2010.00170.x. PMID 21504532. S2CID 7690884.
^ Julian GE, Cahan S (August 1999). "Undertaking specialization in the desert leaf-cutter ant Acromyrmex versicolor". Animal Behaviour. 58 (2): 437–442. doi:10.1006/anbe.1999.1184. PMID 10458895. S2CID 23845331.
^ López-Riquelme GO, Malo EA, Cruz-López L, Fanjul-Moles ML (2006). "Antennal olfactory sensitivity in response to task-related odours of three castes of the ant Atta mexicana (hymenoptera: formicidae)". Physiological Entomology. 31 (4): 353–360. doi:10.1111/j.1365-3032.2006.00526.x. S2CID 84890901.
^ Choe DH, Millar JG, Rust MK (May 2009). "Chemical signals associated with life inhibit necrophoresis in Argentine ants". Proceedings of the National Academy of Sciences of the United States of America. 106 (20): 8251–8255. Bibcode:2009PNAS..106.8251C. doi:10.1073/pnas.0901270106. PMC 2688878. PMID 19416815.
^ Tschinkel WR (2004). "The nest architecture of the Florida harvester ant, Pogonomyrmex badius". Journal of Insect Science. 4 (21): 21. doi:10.1093/jis/4.1.21. PMC 528881. PMID 15861237.
^ Peeters C, Hölldobler B, Moffett M, Musthak Ali TM (1994). ""Wall-papering" and elaborate nest architecture in the ponerine ant Harpegnathos saltator". Insectes Sociaux. 41 (2): 211–218. doi:10.1007/BF01240479. S2CID 41870857.
^ Maschwitz U, Moog J (December 2000). "Communal peeing: a new mode of flood control in ants". Die Naturwissenschaften. 87 (12): 563–565. Bibcode:2000NW.....87..563M. doi:10.1007/s001140050780. PMID 11198200. S2CID 7482935.
^ Nielsen MG, Christian KA (May 2007). "The mangrove ant, Camponotus anderseni, switches to anaerobic respiration in response to elevated CO2 levels". Journal of Insect Physiology. 53 (5): 505–508. doi:10.1016/j.jinsphys.2007.02.002. PMID 17382956.
^ Franks NR, Richardson T (January 2006). "Teaching in tandem-running ants". Nature. 439 (7073): 153. Bibcode:2006Natur.439..153F. doi:10.1038/439153a. PMID 16407943. S2CID 4416276.
^ Ravary F, Lecoutey E, Kaminski G, Châline N, Jaisson P (August 2007). "Individual experience alone can generate lasting division of labor in ants". Current Biology. 17 (15): 1308–1312. Bibcode:2007CBio...17.1308R. doi:10.1016/j.cub.2007.06.047. PMID 17629482. S2CID 13273984.
^ Franks NR, Hooper J, Webb C, Dornhaus A (June 2005). "Tomb evaders: house-hunting hygiene in ants". Biology Letters. 1 (2): 190–192. doi:10.1098/rsbl.2005.0302. PMC 1626204. PMID 17148163.
^ Hölldobler & Wilson (1990), p. 573
^ Robson SK, Kohout RJ (2005). "Evolution of nest-weaving behaviour in arboreal nesting ants of the genus Polyrhachis Fr. Smith (Hymenoptera: Formicidae)". Australian Journal of Entomology. 44 (2): 164–169. doi:10.1111/j.1440-6055.2005.00462.x.
^ Frouz J (2000). "The Effect of Nest Moisture on Daily Temperature Regime in the Nests of Formica polyctena Wood Ants". Insectes Sociaux. 47 (3): 229–235. doi:10.1007/PL00001708. S2CID 955282.
^ Kadochová, Štěpánka; Frouz, Jan (2013). "Thermoregulation strategies in ants in comparison to other social insects, with a focus on red wood ants ( Formica rufa group)". F1000Research. 2: 280. doi:10.12688/f1000research.2-280.v2. ISSN 2046-1402. PMC 3962001. PMID 24715967.
^ Carpintero S, Reyes-López J, de Reynac LA (2004). "Impact of human dwellings on the distribution of the exotic Argentine ant: a case study in the Doñana National Park, Spain". Biological Conservation. 115 (2): 279–289. Bibcode:2004BCons.115..279C. doi:10.1016/S0006-3207(03)00147-2.
^ Friedrich R, Philpott SM (June 2009). "Nest-site limitation and nesting resources of ants (Hymenoptera: Formicidae) in urban green spaces". Environmental Entomology. 38 (3): 600–607. doi:10.1603/022.038.0311. PMID 19508768. S2CID 20555077.
^ Anderson KE, Russell JA, Moreau CS, Kautz S, Sullam KE, Hu Y, Basinger U, Mott BM, Buck N, Wheeler DE (May 2012). "Highly similar microbial communities are shared among related and trophically similar ant species". Molecular Ecology. 21 (9): 2282–2296. Bibcode:2012MolEc..21.2282A. doi:10.1111/j.1365-294x.2011.05464.x. PMID 22276952. S2CID 32534515.
^ Feldhaar H, Straka J, Krischke M, Berthold K, Stoll S, Mueller MJ, Gross R (October 2007). "Nutritional upgrading for omnivorous carpenter ants by the endosymbiont Blochmannia". BMC Biology. 5: 48. doi:10.1186/1741-7007-5-48. PMC 2206011. PMID 17971224.
^ Russell JA, Moreau CS, Goldman-Huertas B, Fujiwara M, Lohman DJ, Pierce NE (December 2009). "Bacterial gut symbionts are tightly linked with the evolution of herbivory in ants". Proceedings of the National Academy of Sciences of the United States of America. 106 (50): 21236–21241. Bibcode:2009PNAS..10621236R. doi:10.1073/pnas.0907926106. PMC 2785723. PMID 19948964.
^ Schultz TR (1999). "Ants, plants and antibiotics" (PDF). Nature. 398 (6730): 747–748. Bibcode:1999Natur.398..747S. doi:10.1038/19619. S2CID 5167611. Archived (PDF) from the original on 9 October 2022.
^ a b c Carrol CR, Janzen DH (1973). "Ecology of foraging by ants". Annual Review of Ecology and Systematics. 4: 231–257. doi:10.1146/annurev.es.04.110173.001311.
^ Wittlinger M, Wehner R, Wolf H (June 2006). "The ant odometer: stepping on stilts and stumps" (PDF). Science. 312 (5782): 1965–1967. Bibcode:2006Sci...312.1965W. doi:10.1126/science.1126912. PMID 16809544. S2CID 15162376. Archived from the original (PDF) on 28 July 2011.
^ Ronacher B, Werner R (1995). "Desert ants Cataglyphis fortis use self-induced optic flow to measure distances travelled" (PDF). Journal of Comparative Physiology A. 177. doi:10.1007/BF00243395. S2CID 4625001. Archived from the original (PDF) on 27 July 2011. Retrieved 7 June 2011.
^ Wehner R (August 2003). "Desert ant navigation: how miniature brains solve complex tasks" (PDF). Journal of Comparative Physiology A. 189 (8): 579–588. doi:10.1007/s00359-003-0431-1. PMID 12879352. S2CID 4571290. Archived from the original (PDF) on 7 July 2011. Retrieved 7 September 2010.
^ Sommer S, Wehner R (January 2004). "The ant's estimation of distance travelled: experiments with desert ants, Cataglyphis fortis" (PDF). Journal of Comparative Physiology A. 190 (1): 1–6. doi:10.1007/s00359-003-0465-4. PMID 14614570. S2CID 23280914.
^ Åkesson S, Wehner R (2002). "Visual navigation in desert ants Cataglyphis fortis: are snapshots coupled to a celestial system of reference?" (PDF). Journal of Experimental Biology. 205 (14): 1971–1978. doi:10.1242/jeb.205.14.1971. PMID 12089203.
^ Steck K, Hansson BS, Knaden M (February 2009). "Smells like home: Desert ants, Cataglyphis fortis, use olfactory landmarks to pinpoint the nest". Frontiers in Zoology. 6: 5. doi:10.1186/1742-9994-6-5. PMC 2651142. PMID 19250516.
^ Seidl T, Wehner R (September 2006). "Visual and tactile learning of ground structures in desert ants". The Journal of Experimental Biology. 209 (Pt 17): 3336–3344. doi:10.1242/jeb.02364. PMID 16916970. S2CID 9642888.
^ Banks AN, Srygley RB (2003). "Orientation by magnetic field in leaf-cutter ants, Atta colombica (Hymenoptera: Formicidae)". Ethology. 109 (10): 835–846. Bibcode:2003Ethol.109..835B. doi:10.1046/j.0179-1613.2003.00927.x.
^ Fukushi T (June 2001). "Homing in wood ants, Formica japonica: use of the skyline panorama". The Journal of Experimental Biology. 204 (Pt 12): 2063–2072. doi:10.1242/jeb.204.12.2063. PMID 11441048.
^ Wehner R, Menzel R (April 1969). "Homing in the ant Cataglyphis bicolor". Science. 164 (3876): 192–194. Bibcode:1969Sci...164..192W. doi:10.1126/science.164.3876.192. PMID 5774195. S2CID 41669795.
^ Chapman RF (1998). The Insects: Structure and Function (4th ed.). Cambridge University Press. pp. 600. ISBN 978-0-521-57890-5.
^ Delsuc F (November 2003). "Army ants trapped by their evolutionary history". PLOS Biology. 1 (2): E37. doi:10.1371/journal.pbio.0000037. PMC 261877. PMID 14624241.
^ Baroni-Urbani C, Boyan GS, Blarer A, Billen J, Musthak Ali TM (1994). "A novel mechanism for jumping in the Indian ant Harpegnathos saltator (Jerdon) (Formicidae, Ponerinae)". Experientia. 50: 63–71. doi:10.1007/BF01992052. S2CID 42304237.
^ Yanoviak SP, Dudley R, Kaspari M (February 2005). "Directed aerial descent in canopy ants" (PDF). Nature. 433 (7026): 624–626. Bibcode:2005Natur.433..624Y. doi:10.1038/nature03254. PMID 15703745. S2CID 4368995. Archived from the original (PDF) on 16 June 2007.
^ Mlot NJ, Tovey CA, Hu DL (May 2011). "Fire ants self-assemble into waterproof rafts to survive floods". Proceedings of the National Academy of Sciences of the United States of America. 108 (19): 7669–7673. Bibcode:2011PNAS..108.7669M. doi:10.1073/pnas.1016658108. PMC 3093451. PMID 21518911.
^ Morrison LW (1998). "A review of Bahamian ant (Hymenoptera: Formicidae) biogeography". Journal of Biogeography. 25 (3): 561–571. Bibcode:1998JBiog..25..561M. doi:10.1046/j.1365-2699.1998.2530561.x. S2CID 84923599.
^ Clay RE, Andersen AN (1996). "Ant fauna of a mangrove community in the Australian seasonal tropics, with particular reference to zonation". Australian Journal of Zoology. 44 (5): 521–533. doi:10.1071/ZO9960521.
^ Crosland MW, Crozier RH, Jefferson E (1988). "Aspects of the biology of the primitive ant genus Myrmecia F. (Hymenoptera: Formicidae)". Australian Journal of Entomology. 27 (4): 305–309. doi:10.1111/j.1440-6055.1988.tb01179.x.
^ Diehl E, Junqueira LK, Berti-Filho E (August 2005). "Ant and termite mound coinhabitants in the wetlands of Santo Antonio da Patrulha, Rio Grande do Sul, Brazil". Brazilian Journal of Biology. 65 (3): 431–437. doi:10.1590/S1519-69842005000300008. PMID 16341421.
^ Achenbach A, Foitzik S (April 2009). "First evidence for slave rebellion: enslaved ant workers systematically kill the brood of their social parasite protomognathus americanus". Evolution; International Journal of Organic Evolution. 63 (4): 1068–1075. doi:10.1111/j.1558-5646.2009.00591.x. PMID 19243573. S2CID 9546342. See also New Scientist, April 9, 2009
^ Henderson G, Andersen JF, Phillips JK, Jeanne RL (July 1990). "Internest aggression and identification of possible nestmate discrimination pheromones in polygynous antFormica montana". Journal of Chemical Ecology. 16 (7): 2217–2228. Bibcode:1990JCEco..16.2217H. doi:10.1007/BF01026932. PMID 24264088. S2CID 22878651.
^ Ward PS (1996). "A new workerless social parasite in the ant genus Pseudomyrmex (Hymenoptera: Formicidae), with a discussion of the origin of social parasitism in ants". Systematic Entomology. 21 (3): 253–263. Bibcode:1996SysEn..21..253W. doi:10.1046/j.1365-3113.1996.d01-12.x. S2CID 84198690.
^ Taylor RW (1968). "The Australian workerless inquiline ant, Strumigenys xenos Brown (Hymenoptera-Formicidae) recorded from New Zealand". New Zealand Entomologist. 4 (1): 47–49. Bibcode:1968NZEnt...4...47T. doi:10.1080/00779962.1968.9722888. S2CID 83791596.
^ Hölldobler & Wilson (1990), pp. 436–448
^ Fournier D, Estoup A, Orivel J, Foucaud J, Jourdan H, Le Breton J, Keller L (June 2005). "Clonal reproduction by males and females in the little fire ant" (PDF). Nature. 435 (7046): 1230–1234. Bibcode:2005Natur.435.1230F. doi:10.1038/nature03705. PMID 15988525. S2CID 1188960.
^ Reiskind J (1977). "Ant-mimicry in Panamanian clubionid and salticid spiders (Araneae: Clubionidae, Salticidae)". Biotropica. 9 (1): 1–8. Bibcode:1977Biotr...9....1R. doi:10.2307/2387854. JSTOR 2387854.
^ Cushing PE (1997). "Myrmecomorphy and myrmecophily in spiders: A Review" (PDF). The Florida Entomologist. 80 (2): 165–193. doi:10.2307/3495552. JSTOR 3495552. Archived (PDF) from the original on 9 October 2022.
^ Styrsky JD, Eubanks MD (January 2007). "Ecological consequences of interactions between ants and honeydew-producing insects". Proceedings. Biological Sciences. 274 (1607): 151–164. doi:10.1098/rspb.2006.3701. PMC 1685857. PMID 17148245.
^ Jahn GC, Beardsley JW (1996). "Effects of Pheidole megacephala (Hymenoptera: Formicidae) on survival and dispersal of Dysmicoccus neobrevipes (Homoptera: Pseudococcidae)". Journal of Economic Entomology. 89 (5): 1124–1129. doi:10.1093/jee/89.5.1124.
^ DeVries PJ (1992). "Singing caterpillars, ants and symbiosis". Scientific American. 267 (4): 76–82. Bibcode:1992SciAm.267d..76D. doi:10.1038/scientificamerican1092-76.
^ Elfferich NW (1998). "Is the larval and imaginal signalling of Lycaenidae and other Lepidoptera related to communication with ants". Deinsea. 4 (1).
^ Pierce NE, Braby MF, Heath A, Lohman DJ, Mathew J, Rand DB, Travassos MA (2002). "The ecology and evolution of ant association in the Lycaenidae (Lepidoptera)". Annual Review of Entomology. 47: 733–771. doi:10.1146/annurev.ento.47.091201.145257. PMID 11729090.
^ Kautz S, Rubin BE, Russell JA, Moreau CS (January 2013). "Surveying the microbiome of ants: comparing 454 pyrosequencing with traditional methods to uncover bacterial diversity". Applied and Environmental Microbiology. 79 (2): 525–534. Bibcode:2013ApEnM..79..525K. doi:10.1128/AEM.03107-12. PMC 3553759. PMID 23124239.
^ Neuvonen MM, Tamarit D, Näslund K, Liebig J, Feldhaar H, Moran NA, Guy L, Andersson SG (December 2016). "The genome of Rhizobiales bacteria in predatory ants reveals urease gene functions but no genes for nitrogen fixation". Scientific Reports. 6 (1): 39197. Bibcode:2016NatSR...639197N. doi:10.1038/srep39197. PMC 5156944. PMID 27976703.
^ Dejean A, Solano PJ, Ayroles J, Corbara B, Orivel J (April 2005). "Insect behaviour: arboreal ants build traps to capture prey". Nature. 434 (7036): 973. Bibcode:2005Natur.434..973D. doi:10.1038/434973a. PMID 15846335. S2CID 4428574.
^ Frederickson ME, Gordon DM (April 2007). "The devil to pay: a cost of mutualism with Myrmelachista schumanni ants in 'devil's gardens' is increased herbivory on Duroia hirsuta trees". Proceedings. Biological Sciences. 274 (1613): 1117–23. doi:10.1098/rspb.2006.0415. PMC 2124481. PMID 17301016.
^ Peakall R, Beattie AJ, James SH (October 1987). "Pseudocopulation of an orchid by male ants: a test of two hypotheses accounting for the rarity of ant pollination". Oecologia. 73 (4): 522–524. Bibcode:1987Oecol..73..522P. doi:10.1007/BF00379410. PMID 28311968. S2CID 3195610.
^ Beattie AJ, Turnbull C, R B K, E G W (1984). "Ant Inhibition of Pollen Function: A Possible Reason Why Ant Pollination is Rare". American Journal of Botany. 71 (3): 421–426. doi:10.2307/2443499. JSTOR 2443499.
^ New, Tim R. (2017). "Classic Themes: Ants, Plants and Fungi". Mutualisms and Insect Conservation. Springer International Publishing. pp. 63–103. doi:10.1007/978-3-319-58292-4_4. ISBN 9783319582917.
^ Katayama N, Suzuki N (2004). "Role of extrafloral nectaries of Vicia faba in attraction of ants and herbivore exclusion by ants". Entomological Science. 7 (2): 119–124. doi:10.1111/j.1479-8298.2004.00057.x. hdl:10252/00005880. S2CID 85428729.
^ Fischer RC, Wanek W, Richter A, Mayer V (2003). "Do ants feed plants? A 15N labelling study of nitrogen fluxes from ants to plants in the mutualism of Pheidole and Piper". Journal of Ecology. 91 (1): 126–134. Bibcode:2003JEcol..91..126F. doi:10.1046/j.1365-2745.2003.00747.x.
^ Chomicki G, Renner SS (November 2016). "Obligate plant farming by a specialized ant". Nature Plants. 2 (12): 16181. doi:10.1038/nplants.2016.181. PMID 27869787. S2CID 23748032.
^ Chomicki G, Janda M, Renner SS (March 2017). "The assembly of ant-farmed gardens: mutualism specialization following host broadening". Proceedings. Biological Sciences. 284 (1850): 20161759. doi:10.1098/rspb.2016.1759. PMC 5360912. PMID 28298344.
^ Fiala B, Maschwitz U, Pong TY, Helbig AJ (June 1989). "Studies of a South East Asian ant-plant association: protection of Macaranga trees by Crematogaster borneensis" (PDF). Oecologia. 79 (4): 463–470. Bibcode:1989Oecol..79..463F. doi:10.1007/bf00378662. PMID 28313479. S2CID 21112371.
^ a b Lengyel S, Gove AD, Latimer AM, Majer JD, Dunn RR (2010). "Convergent evolution of seed dispersal by ants, and phylogeny and biogeography in flowering plants: A global survey". Perspectives in Plant Ecology, Evolution and Systematics. 12: 43–55. doi:10.1016/j.ppees.2009.08.001.
^ Giladi I (2006). "Choosing benefits or partners: a review of the evidence for the evolution of myrmecochory". Oikos. 112 (3): 481–492. Bibcode:2006Oikos.112..481G. CiteSeerX 10.1.1.530.1306. doi:10.1111/j.0030-1299.2006.14258.x.
^ Hanzawa FM, Beattie AJ, Culver DC (1988). "Directed dispersal: demographic analysis of an ant-seed mutualism". American Naturalist. 131 (1): 1–13. doi:10.1086/284769. S2CID 85317649.
^ Auld TD (1996). "Ecology of the Fabaceae in the Sydney region: fire, ants and the soil seedbank". Cunninghamia. 4 (22).
^ Fischer RC, Ölzant SM, Wanek W, Mayer V (2005). "The fate of Corydalis cava elaiosomes within an ant colony of Myrmica rubra: elaiosomes are preferentially fed to larvae". Insectes Sociaux. 52 (1): 55–62. doi:10.1007/s00040-004-0773-x. S2CID 21974767.
^ Frouz, Jan (1997). "The effect of wood ants (Formica polyctena Foerst) on the transformation of phosphorus in a spruce plantation". Pedobiologia. 41 (5): 437–447. doi:10.1016/S0031-4056(24)00314-7.
^ Frouz, J; Jílková, V (2008). "The effect of ants on soil properties and processes (Hymenoptera: Formicidae)". Myrmecological News. 1: 191–199.
^ Frouz, Jan; Jílková, Veronika; Sorvari, Jouni (2016). "Contribution of wood ants to nutrient cycling and ecosystem function". In Robinson, Elva J. H.; Stockan, Jenni A. (eds.). Wood Ant Ecology and Conservation. Ecology, Biodiversity and Conservation. Cambridge: Cambridge University Press. pp. 207–220. ISBN 978-1-107-04833-1. Retrieved 12 July 2021.
^ Hughes L, Westoby M (1992). "Capitula on stick insect eggs and elaiosomes on seeds: convergent adaptations for burial by ants". Functional Ecology. 6 (6): 642–648. Bibcode:1992FuEco...6..642.. doi:10.2307/2389958. JSTOR 2389958.
^ Quinet Y, Tekule N, de Biseau JC (2005). "Behavioural Interactions Between Crematogaster brevispinosa rochai Forel (Hymenoptera: Formicidae) and Two Nasutitermes Species (Isoptera: Termitidae)". Journal of Insect Behavior. 18 (1): 1–17. Bibcode:2005JIBeh..18....1Q. doi:10.1007/s10905-005-9343-y. S2CID 33487814.
^ Jeanne RL (1972). "Social biology of the neotropical wasp Mischocyttarus drewseni". Bull. Mus. Comp. Zool. 144: 63–150.
^ Jeanne R (July 1995). "Foraging in Social Wasps: Agelaia lacks recruitment to food (Hymenoptera: Vespidae)". Journal of the Kansas Entomological Society.
^ a b Sivinski J, Marshall S, Petersson E (1999). "Kleptoparasitism and phoresy in the Diptera" (PDF). Florida Entomologist. 82 (2): 179–197. doi:10.2307/3496570. JSTOR 3496570. Archived (PDF) from the original on 9 October 2022.
^ Schaechter E (2000). "Some weird and wonderful fungi". Microbiology Today. 27 (3): 116–117.
^ Andersen SB, Gerritsma S, Yusah KM, Mayntz D, Hywel-Jones NL, Billen J, Boomsma JJ, Hughes DP (September 2009). "The life of a dead ant: the expression of an adaptive extended phenotype" (PDF). The American Naturalist. 174 (3): 424–433. doi:10.1086/603640. hdl:11370/e6374602-b2a0-496c-b78e-774b34fb152b. PMID 19627240. S2CID 31283817.
^ Wojcik DP (1989). "Behavioral interactions between ants and their parasites". The Florida Entomologist. 72 (1): 43–451. doi:10.2307/3494966. JSTOR 3494966. Archived from the original (PDF) on 9 March 2021. Retrieved 25 October 2017.
^ Poinar G, Yanoviak SP (2008). "Myrmeconema neotropicum n. g., n. sp., a new tetradonematid nematode parasitising South American populations of Cephalotes atratus (Hymenoptera: Formicidae), with the discovery of an apparent parasite-induced host morph". Systematic Parasitology. 69 (2): 145–153. doi:10.1007/s11230-007-9125-3. PMID 18038201. S2CID 8473071.
^ Stoldt M, Klein L, Beros S, Butter F, Jongepier E, Feldmeyer B, Foitzik S (January 2021). "Parasite Presence Induces Gene Expression Changes in an Ant Host Related to Immunity and Longevity". Genes. 12 (1): 202118. doi:10.1098/rsos.202118. PMC 8131941. PMID 34017599.
^ Caldwell JP (1996). "The evolution of myrmecophagy and its correlates in poison frogs (Family Dendrobatidae)". Journal of Zoology. 240 (1): 75–101. doi:10.1111/j.1469-7998.1996.tb05487.x.
^ Willis E, Oniki Y (1978). "Birds and Army Ants". Annual Review of Ecology and Systematics. 9: 243–263. doi:10.1146/annurev.es.09.110178.001331.
^ Vellely AC (2001). "Foraging at army ant swarms by fifty bird species in the highlands of Costa Rica" (PDF). Ornitologia Neotropical. 12: 271–275. Archived (PDF) from the original on 9 October 2022. Retrieved 8 June 2008.
^ Inzunza ER, Martínez-Leyva JE, Valenzuela-González JE (2015). "Doves kleptoparasitize ants". The Southwestern Naturalist. 60 (1): 103–106. doi:10.1894/msh-03.1. S2CID 85633598.
^ Wrege PH, Wikelski M, Mandel JT, Rassweiler T, Couzin ID (2005). "Antbirds parasitize foraging army ants". Ecology. 86 (3): 555–559. Bibcode:2005Ecol...86..555W. doi:10.1890/04-1133.
^ Swenson JE, Jansson A, Riig R, Sandegren R (1999). "Bears and ants: myrmecophagy by brown bears in central Scandinavia". Canadian Journal of Zoology. 77 (4): 551–561. doi:10.1139/z99-004.
^ Hölldobler & Wilson (1990), pp. 619–629
^ Gottrup F, Leaper D (2004). "Wound healing: Historical aspects" (PDF). EWMA Journal. 4 (2): 5. Archived from the original (PDF) on 16 June 2007.
^ Gudger EW (1925). "Stitching wounds with the mandibles of ants and beetles". Journal of the American Medical Association. 84 (24): 1861–1864. doi:10.1001/jama.1925.02660500069048.
^ Sapolsky RM (2001). A Primate's Memoir: A Neuroscientist's Unconventional Life Among the Baboons. Simon and Schuster. pp. 156. ISBN 978-0-7432-0241-1.
^ Wheeler WM (1910). Ants: Their Structure, Development and Behavior . Columbia University Biological Series. Vol. 9. Columbia University Press. p. 10. doi:10.5962/bhl.title.1937. ISBN 978-0-231-00121-2. LCCN 10008253//r88. OCLC 560205.
^ Haddad Junior V, Cardoso JL, Moraes RH (2005). "Description of an injury in a human caused by a false tocandira (Dinoponera gigantea, Perty, 1833) with a revision on folkloric, pharmacological and clinical aspects of the giant ants of the genera Paraponera and Dinoponera (sub-family Ponerinae)". Revista do Instituto de Medicina Tropical de Sao Paulo. 47 (4): 235–238. doi:10.1590/S0036-46652005000400012. hdl:11449/30504. PMID 16138209.
^ McGain F, Winkel KD (August 2002). "Ant sting mortality in Australia". Toxicon. 40 (8): 1095–1100. doi:10.1016/S0041-0101(02)00097-1. PMID 12165310.
^ Downes D, Laird SA (1999). "Innovative mechanisms for sharing benefits of biodiversity and related knowledge" (PDF). The Center for International Environmental Law. Archived (PDF) from the original on 23 April 2008. Retrieved 8 June 2008.
^ Cheney RH, Scholtz E (1963). "Rooibos tea, a South African contribution to world beverages". Economic Botany. 17 (3): 186–194. doi:10.1007/BF02859435. S2CID 37728834.
^ Chapman RE, Bourke AF (2001). "The influence of sociality on the conservation biology of social insects". Ecology Letters. 4 (6): 650–662. Bibcode:2001EcolL...4..650C. doi:10.1046/j.1461-0248.2001.00253.x. S2CID 86796899.
^ Cruz-Labana, J. D.; Tarango-Arámbula, L. A.; Alcántara-Carbajal, J. L.; Pimentel-López, J.; Ugalde-Lezama, S.; Ramírez-Valverde, G.; Méndez-Gallegos, S. J. (2014). "Habitat use by the "Escamolera" ant (Liometopum apiculatum Mayr) in central Mexico". Agrociencia. 48 (6): 569–582. ISSN 1405-3195.
^ DeFoliart GR (1999). "Insects as food: why the western attitude is important". Annual Review of Entomology. 44: 21–50. doi:10.1146/annurev.ento.44.1.21. PMID 9990715.
^ Bingham, C.T. (1903). Fauna of British India. Hymenoptera. Volume 2. London: Taylor and Francis. p. 311.
^ a b Bequaert J (1921). "Insects as food: How they have augmented the food supply of mankind in early and recent times". Natural History Journal. 21: 191–200.
^ a b "Pest Notes: Ants (Publication 7411)". University of California Agriculture and Natural Resources. 2007. Retrieved 5 June 2008.
^ Lubbock J (1881). "Observations on ants, bees, and wasps. IX. Color of flowers as an attraction to bees: Experiments and considerations thereon". J. Linn. Soc. Lond. (Zool.). 16 (90): 110–112. doi:10.1111/j.1096-3642.1882.tb02275.x.
^ Stadler B, Dixon AF (2008). Mutualism: Ants and their insect partners. Cambridge University Press. ISBN 978-0-521-86035-2.
^ Kennedy CH (1951). "Myrmecological technique. IV. Collecting ants by rearing pupae". The Ohio Journal of Science. 51 (1): 17–20. hdl:1811/3802.
^ Wojcik DP, Burges RJ, Blanton CM, Focks DA (2000). "An improved and quantified technique for marking individual fire ants (Hymenoptera: Formicidae)". The Florida Entomologist. 83 (1): 74–78. doi:10.2307/3496231. JSTOR 3496231.
^ Dicke E, Byde A, Cliff D, Layzell P (2004). Ispeert AJ, Murata M, Wakamiya N (eds.). "An ant-inspired technique for storage area network design". Proceedings of Biologically Inspired Approaches to Advanced Information Technology: First International Workshop, BioADIT 2004 LNCS 3141: 364–379.
^ US granted 5803014, Guri A, "Habitat media for ants and other invertebrates", issued 8 September 1998, assigned to Plant Cell Technology Inc
^ "The Ant, The Ants". Quran. Vol. Surah 27. pp. 18–19. Archived from the original on 1 January 2007.
^ Bukhari S. "Beginning of Creation". Sunnah. Vol. 4 Book 54. Archived from the original on 18 August 2000.
^ See wikisource:Bible (World English)/Proverbs#30:25
^ Mentioned once in the Quran, Muhammad Asad translates the verse as following: till, when they came upon a valley [full] of ants, and an ant exclaimed: "O you ants! Get into your dwellings, lest Solomon and his hosts crush you without [even] being aware [of you]! (27:18)"
^ Deen MY (1990). "Islamic Environmental Ethics, Law, and Society" (PDF). In Engel JR, Engel JG (eds.). Ethics of Environment and Development. Bellhaven Press, London. Archived from the original (PDF) on 14 July 2011.
^ Balee WL (2000). "Antiquity of traditional ethnobiological knowledge in Amazonia: The Tupi-Guarani family and time". Ethnohistory. 47 (2): 399–422. doi:10.1215/00141801-47-2-399. S2CID 162813070.
^ Cesard N, Deturche J, Erikson P (2003). "Les Insectes dans les pratiques médicinales et rituelles d'Amazonie indigène". In Motte-Florac E, Thomas JM (eds.). Les insectes dans la tradition orale (in French). Peeters-Selaf, Paris. pp. 395–406.
^ Schmidt RJ (May 1985). "The super-nettles. A dermatologist's guide to ants in the plants". International Journal of Dermatology. 24 (4): 204–210. doi:10.1111/j.1365-4362.1985.tb05760.x. PMID 3891647. S2CID 73875767.
^ Servius, Commentary on Virgil's Aeneid 4.402; Smith 1873, s.v. Myrmex
^ Twain M (1880). "22 The Black Forest and Its Treasures". A Tramp Abroad. New York: Oxford University Press. ISBN 978-0-19-510137-9. Retrieved 13 December 2015.
^ Winthrop-Young, Geoffrey (2021). "A Green Hell Makes Better Germans: Carl Stephenson's "Leiningen" and the Almost Aryan Countertextual Army Ants". The Germanic Review: Literature, Culture, Theory. 96 (4): 339–356. doi:10.1080/00168890.2021.1977228. ISSN 0016-8890. S2CID 245125375.
^ Wilson, EO (25 January 2010). "Trailhead". The New Yorker. pp. 56–62.
^ Parent A (August 2003). "Auguste Forel on ants and neurology". The Canadian Journal of Neurological Sciences. 30 (3): 284–291. doi:10.1017/s0317167100002754. PMID 12945958.
^ "1992 Excellence in Software Awards Winners". Software & Information Industry Association. Archived from the original on 11 June 2009. Retrieved 3 April 2008.
^ Sharkey AJC (2006). "Robots, insects and swarm intelligence". Artificial Intelligence Review. 26 (4): 255–268. doi:10.1007/s10462-007-9057-y. S2CID 321326.
Cited texts
Borror DJ, Triplehorn CA, Delong DM (1989). Introduction to the Study of Insects (6th ed.). Saunders College Publishing. ISBN 978-0-03-025397-3.
Hölldobler B, Wilson EO (1990). The Ants. Harvard University Press. ISBN 978-0-674-04075-5.
Further reading
Mike Snider (8 January 2022). "When the trees where these ants live were damaged, they made some DIY home repairs". USA Today.
Bolton, Barry (1995). A New General Catalogue of the Ants of the World. Harvard University Press. ISBN 978-0-674-61514-4.
Hölldobler B, Wilson EO (1998). Journey to the Ants: A Story of Scientific Exploration. Belknap Press. ISBN 978-0-674-48526-6.
Hölldobler B, Wilson EO (2009). The Superorganism: The Beauty, Elegance and Strangeness of Insect Societies. Norton & Co. ISBN 978-0-393-06704-0.
External links
Wikiquote has quotations related to Ant.
Wikimedia Commons has media related to Formicidae.
Wikispecies has information related to Formicidae.
Wilson, Andrew (1878). "Ant" . Encyclopædia Britannica. Vol. II (9th ed.). pp. 94–100.
AntWeb from The California Academy of Sciences
AntWiki – Bringing Ants to the World
Ant Species Fact Sheets from the National Pest Management Association on Argentine, Carpenter, Pharaoh, Odorous, and other ant species
Ant Genera of the World – distribution maps
The super-nettles. A dermatologist's guide to ants-in-the-plants
vteExtant Hymenopteran families
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Subclass: Pterygota
Infraclass: Neoptera
Superorder: Holometabola
SymphytaTenthredinoidea
Argidae (argid sawflies)
Blasticotomidae (fern sawflies)
Cimbicidae (cimbicid sawflies)
Diprionidae (conifer sawflies)
Pergidae (pergid sawflies)
Tenthredinidae (common sawflies)
XyeloideaXyelidaePamphilioidea
Megalodontesidae
Pamphiliidae (web-spinning sawflies)
Siricoidea
Anaxyelidae (cedar wood wasps)
Siricidae (horntails)
Xiphydrioidea
Xiphydriidae (wood wasps)
Cephoidea
Cephidae (stem sawflies)
Orussoidea
Orussidae (parasitic wood wasps)
ApocritaParasiticaIchneumonoidea
Braconidae (braconids)
Ichneumonidae (ichneumon wasps)
Ceraphronoidea
Ceraphronidae
Megaspilidae
ProctotrupomorphaPlatygastroidea
Geoscelionidae
Janzenellidae
Neuroscelionidae
Nixoniidae
Platygastridae
Scelionidae
Sparasionidae
Cynipoidea
Austrocynipidae
Cynipidae (gall wasps)
Diplolepididae
Figitidae
Ibaliidae
Liopteridae
Proctotrupoidea (s.str.)
Proctorenyxidae
Roproniidae
Heloridae
Pelecinidae
Peradeniidae
Proctotrupidae
Vanhorniidae
Diaprioidea
Austroniidae
Diapriidae
Ismaridae
Maamingidae
Monomachidae
Mymarommatoidea
Mymarommatidae
Chalcidoidea(chalcid wasps)
Agaonidae (fig wasps)
Aphelinidae
Azotidae
Baeomorphidae
Boucekiidae
Calesidae
Ceidae
Cerocephalidae
Chalcedectidae
Chalcididae
Chrysolampidae
Cleonymidae
Coelocybidae
Cynipencyrtidae
Diparidae
Encyrtidae
Epichrysomallidae
Eucharitidae
Eulophidae
Eunotidae
Eupelmidae
Eurytomidae
Eutrichosomatidae
Herbertiidae
Hetreulophidae
Heydeniidae
Idioporidae
Leucospidae
Lyciscidae
Macromesidae
Megastigmidae
Melanosomellidae
Metapelmatidae
Moranilidae
Mymaridae (fairyflies)
Neanastatidae
Neodiparidae
Ooderidae
Ormyridae
Pelecinellidae
Perilampidae
Pirenidae
Pteromalidae
Signiphoridae
Spalangiidae
Systasidae
Tanaostigmatidae
Tetracampidae
Torymidae
Trichogrammatidae
Evanioidea
Aulacidae
Evaniidae (ensign wasps)
Gasteruptiidae
Stephanoidea
Stephanidae
Megalyroidea
Megalyridae
Trigonaloidea
Trigonalidae
AculeataChrysidoidea
Bethylidae
Chrysididae (cuckoo wasps)
Dryinidae
Embolemidae
Plumariidae
Sclerogibbidae
Scolebythidae
Vespoidea
Rhopalosomatidae (rhopalosomatid wasps)
Vespidae (paper wasps, potter wasps, pollen wasps, yellowjackets, hornets)
Tiphioidea
Bradynobaenidae
Tiphiidae (tiphiid wasps)
Thynnoidea
Chyphotidae
Thynnidae (flower wasps)
Sierolomorphidae (sierolomorphid wasps)
Pompiloidea
Mutillidae (velvet ants)
Myrmosidae
Pompilidae (spider wasps)
Sapygidae (sapygid, or club-horned wasps)
Scolioidea
Scoliidae (scoliid wasps)
Formicoidea
Formicidae (ants)
ApoideaSpheciformes(sphecoid wasps)
Ammoplanidae
Ampulicidae (cockroach wasps)
Astatidae
Bembicidae
Crabronidae
Mellinidae
Pemphredonidae
Philanthidae
Psenidae
Sphecidae (thread-waisted wasps)
Anthophila(bees)
Andrenidae (mason bees)
Apidae (honey bees, bumblebees, cuckoo bees, carpenter bees, orchid bees, stingless bees)
Colletidae (plasterer bees)
Halictidae (sweat bees)
Megachilidae (mason bees, leafcutter bees)
Melittidae
Stenotritidae
See also: Unicalcarida
vteAnt taxonomy
Kingdom Animalia
Phylum Arthropoda
Class Insecta
Order Hymenoptera
Family Formicidae
SubfamiliesExtant
Agroecomyrmecinae
Amblyoponinae
Aneuretinae
Apomyrminae
Dolichoderinae
Dorylinae
Ectatomminae
Formicinae
Heteroponerinae
Leptanillinae
Martialinae
Myrmeciinae
Myrmicinae
Paraponerinae
Ponerinae
Proceratiinae
Pseudomyrmecinae
Extinct
†Armaniinae
†Brownimeciinae
†Formiciinae
†Haidomyrmecinae
†Sphecomyrminae
Genera
List of ant genera
incertae sedis
vteEusocialityTopics
Evolution of eusociality
Presociality
Social insects
Gamergate
Group selection
Haplodiploidy
Identity in social insects
Kin recognition
Kin selection
Polyethism
Sexual selection in insects
Thelytoky
Worker policing
Groups
Hymenoptera
Ant
Apidae
Crabronidae
Halictidae
Honey bee
Vespidae
Mammalia
Blesmol
Dwarf mongoose
Meerkat
Crustacean
Synalpheus
Thysanoptera
Kladothrips
Hemiptera
Aphididae
Coleoptera
Austroplatypus incompertus
Isoptera
In culture
Bees in mythology
Coalescent
Pioneers, works
Karl von Frisch
The Dancing Bees 1927
Charles Duncan Michener
The Bees of the World 2000
E. O. Wilson
Sociobiology: The New Synthesis 1975
The Ants 1990
Journey to the Ants 1994
vteHuman interactions with insectsAspectsof insectsin cultureIn the arts
Insects in art
Beetlewing
Musca depicta
Arthropods in film
Insects in literature
Insects in music
List of insect-inspired songs
Insects on stamps
In fishing
Fishing bait
Fly fishing
Artificial fly
Fly tying
In medicine
Apitherapy
Apitoxin
Melittin
Spanish fly
Cantharidin
In mythology
Bees in mythology
Cicadas in mythology
Scarab (artifact)
Other aspects
Biomimicry
Cockroach racing
Cricket fighting
Entomological warfare
Entomophagy
Insect farming
Flea circus
Insects in ethics
Insects in religion
Jingzhe
Economicentomology
Beneficial insect
Biological pest control
Beekeeping
Bee pollen
List of crop plants pollinated by bees
Beeswax
Honey
Propolis
Royal jelly
Carmine/Cochineal
Polish
Chitin
Kermes
Sericulture
Silk
Shellac
Model organism
Drosophila melanogaster
Harmfulinsects
Insect bites and stings
Insect sting allergy
Bed bug
Woodworm
Home-stored product entomology
Clothes moth
Pioneers
Jan Swammerdam
Alfred Russel Wallace
Jean-Henri Fabre
Hans Zinsser (Rats, Lice and History)
Lafcadio Hearn (Insect Literature)
Concerns
Bees and toxic chemicals
Colony collapse disorder
Decline in insect populations
Habitat destruction
List of endangered insects
Pesticide
Insecticide
Neonicotinoid
Pesticide toxicity to bees
Categories,templates
Insects and humans
Insecticides
Pesticides
Insects portal
Taxon identifiersFormicidae
Wikidata: Q7386
Wikispecies: Formicidae
ADW: Formicidae
AFD: Formicidae
BOLD: 685
BugGuide: 165
CoL: 623T5
EoL: 699
EPPO: 1FORMF
Fauna Europaea: 11356
Fauna Europaea (new): a9d51445-1393-482b-9ef5-4837fb827590
Fossilworks: 70728
GBIF: 4342
iNaturalist: 47336
IRMNG: 101758
ITIS: 154193
NBN: NBNSYS0000037030
NCBI: 36668
NZOR: cc4a826f-854a-47fa-bb2c-8da6da11ab17
Open Tree of Life: 7376225
Plazi: E378AE58-D311-FFFF-FF78-FB014CA57281
WoRMS: 425369
Authority control databases: National
Spain
France
BnF data
Germany
Israel
United States
Japan
Czech Republic
Retrieved from "https://en.wikipedia.org/w/index.php?title=Ant&oldid=1211959109"
Categories: AntsSymbiosisExtant Albian first appearancesInsects in cultureHidden categories: CS1: long volume valueCS1 French-language sources (fr)Articles with short descriptionShort description matches WikidataFeatured articlesWikipedia pages semi-protected against vandalismWikipedia indefinitely move-protected pagesUse dmy dates from June 2023Articles with 'species' microformatsArticles containing Middle English (1100-1500)-language textArticles containing Old English (ca. 450-1100)-language textArticles containing Low German-language textArticles containing Old Saxon-language textArticles containing German-language textArticles containing Old High German (ca. 750-1050)-language textArticles with text in West Germanic languagesArticles containing Proto-Germanic-language textArticles containing Latin-language textArticles containing Portuguese-language textArticles containing Italian-language textArticles containing Spanish-language textArticles containing Romanian-language textArticles containing French-language textArticles containing Thai-language textCommons category link is on WikidataWikipedia articles incorporating a citation from EB9Taxonbars with 20–24 taxon IDsArticles with BNE identifiersArticles with BNF identifiersArticles with BNFdata identifiersArticles with GND identifiersArticles with J9U identifiersArticles with LCCN identifiersArticles with NDL identifiersArticles with NKC identifiersArticles containing video clips
This page was last edited on 5 March 2024, at 13:13 (UTC).
Text is available under the Creative Commons Attribution-ShareAlike License 4.0;
additional terms may apply. By using this site, you agree to the Terms of Use and Privacy Policy. Wikipedia® is a registered trademark of the Wikimedia Foundation, Inc., a non-profit organization.
Privacy policy
About Wikipedia
Disclaimers
Contact Wikipedia
Code of Conduct
Developers
Statistics
Cookie statement
Mobile view
Toggle limited content width
Ant | Description, Taxonomy, Habitat, Species, Life Cycle, & Facts | Britannica
Search Britannica
Click here to search
Search Britannica
Click here to search
Login
Subscribe
Subscribe
Home
Games & Quizzes
History & Society
Science & Tech
Biographies
Animals & Nature
Geography & Travel
Arts & Culture
Money
Videos
On This Day
One Good Fact
Dictionary
New Articles
History & Society
Lifestyles & Social Issues
Philosophy & Religion
Politics, Law & Government
World History
Science & Tech
Health & Medicine
Science
Technology
Biographies
Browse Biographies
Animals & Nature
Birds, Reptiles & Other Vertebrates
Bugs, Mollusks & Other Invertebrates
Environment
Fossils & Geologic Time
Mammals
Plants
Geography & Travel
Geography & Travel
Arts & Culture
Entertainment & Pop Culture
Literature
Sports & Recreation
Visual Arts
Companions
Demystified
Image Galleries
Infographics
Lists
Podcasts
Spotlights
Summaries
The Forum
Top Questions
#WTFact
100 Women
Britannica Kids
Saving Earth
Space Next 50
Student Center
Home
Games & Quizzes
History & Society
Science & Tech
Biographies
Animals & Nature
Geography & Travel
Arts & Culture
Money
Videos
ant
Table of Contents
ant
Table of Contents
IntroductionPhysical descriptionNatural historyFoodNotable ant behaviors
References & Edit History
Quick Facts & Related Topics
Images & Videos
For Students
ant summary
Quizzes
Creepy Crawlers Quiz
Animal Factoids
Know Your Bugs Quiz
A Is for Animal Quiz
Animal Group Names
Read Next
10 Questions About Insects Answered
Discover
What’s the Difference Between Bison and Buffalo?
Ten Days That Vanished: The Switch to the Gregorian Calendar
The Largest Islands in the World
Who Votes for the Academy Awards?
Celebrating Ramadan
New Seven Wonders of the World
12 Greek Gods and Goddesses
Home
Science
Bugs, Mollusks & Other Invertebrates
Insects
Animals & Nature
ant
insect
Actions
Cite
verifiedCite
While every effort has been made to follow citation style rules, there may be some discrepancies.
Please refer to the appropriate style manual or other sources if you have any questions.
Select Citation Style
MLA
APA
Chicago Manual of Style
Copy Citation
Share
Share
Share to social media
URL
https://www.britannica.com/animal/ant
Give Feedback
External Websites
Feedback
Corrections? Updates? Omissions? Let us know if you have suggestions to improve this article (requires login).
Feedback Type
Select a type (Required)
Factual Correction
Spelling/Grammar Correction
Link Correction
Additional Information
Other
Your Feedback
Submit Feedback
Thank you for your feedback
Our editors will review what you’ve submitted and determine whether to revise the article.
External Websites
Purdue University - College of Agriculture - Extension Entomology - Ants
Live Science - Ants: From the Cool to the Creepy
Animal Corner - Ant
The Australian Museum - Ants: Family Formicidae
A-Z Animals - Ant
Pest World for Kids - Ants
Animal Diversity Web - Formicidae
WebMD - Ants: What to Know
University of Minnesota Extension - Ant
Britannica Websites
Articles from Britannica Encyclopedias for elementary and high school students.
ant - Children's Encyclopedia (Ages 8-11)
ant - Student Encyclopedia (Ages 11 and up)
Please select which sections you would like to print:
Table Of Contents
Cite
verifiedCite
While every effort has been made to follow citation style rules, there may be some discrepancies.
Please refer to the appropriate style manual or other sources if you have any questions.
Select Citation Style
MLA
APA
Chicago Manual of Style
Copy Citation
Share
Share
Share to social media
URL
https://www.britannica.com/animal/ant
Feedback
External Websites
Feedback
Corrections? Updates? Omissions? Let us know if you have suggestions to improve this article (requires login).
Feedback Type
Select a type (Required)
Factual Correction
Spelling/Grammar Correction
Link Correction
Additional Information
Other
Your Feedback
Submit Feedback
Thank you for your feedback
Our editors will review what you’ve submitted and determine whether to revise the article.
External Websites
Purdue University - College of Agriculture - Extension Entomology - Ants
Live Science - Ants: From the Cool to the Creepy
Animal Corner - Ant
The Australian Museum - Ants: Family Formicidae
A-Z Animals - Ant
Pest World for Kids - Ants
Animal Diversity Web - Formicidae
WebMD - Ants: What to Know
University of Minnesota Extension - Ant
Britannica Websites
Articles from Britannica Encyclopedias for elementary and high school students.
ant - Children's Encyclopedia (Ages 8-11)
ant - Student Encyclopedia (Ages 11 and up)
Also known as: Formicidae
Written and fact-checked by
The Editors of Encyclopaedia Britannica
Encyclopaedia Britannica's editors oversee subject areas in which they have extensive knowledge, whether from years of experience gained by working on that content or via study for an advanced degree. They write new content and verify and edit content received from contributors.
The Editors of Encyclopaedia Britannica
Last Updated:
Feb 27, 2024
•
Article History
Table of Contents
bullet ant
See all media
Category:
Animals & Nature
Key People:
E.O. Wilson
Charles Henry Turner
William Morton Wheeler
(Show more)
Related Topics:
leafcutter ant
fire ant
honey ant
harvester ant
army ant
(Show more)
See all related content →
Recent News
Feb. 26, 2024, 4:24 PM ET (Yahoo News)
Homeowner shocked as backyard taken over by ant 'metropolis'
ant, (family Formicidae), any of approximately 10,000 species of insects that are social in habit and live together in organized colonies. Ants occur worldwide but are most numerous, both in numbers and in species, in tropical and subtropical regions. Ants are essential members of the ecosystems they inhabit, and some even serve as keystone species that have a disproportionately large effect on their ecological communities. Some ants are considered pests to humans, and a number are invasive species in areas outside their native ranges.TaxonomyKingdom: AnimaliaPhylum: ArthropodaClass: InsectaOrder: HymenopteraSee also list of ants, bees, and wasps. Physical description black carpenter antBlack carpenter ant (Camponotus pennsylvanicus).(more)Ants range in size from about 2 to 25 mm (about 0.08 to 1 inch). Their color is usually yellow, brown, red, or black. A few genera (e.g., Pheidole of North America) have a metallic luster. Typically, an ant has a large head and a slender, oval abdomen joined to the thorax, or midsection, by a small waist. In all ants there are either one or two finlike extensions running across the thin waist region. The antennae are always elbowed. There are two sets of jaws: the outer pair is used for carrying objects such as food and for digging, and the inner pair is used for chewing. Some species, such as the debilitating bullet ant (Paraponera clavata), have a powerful sting at the tip of the abdomen.
Britannica Quiz
Match the Baby Animal to Its Mama Quiz
Natural history ant bridgeWorker ants form a bridge to cross from one leaf to another.(more)Hear E.O. Wilson talk about his research on how ants determine the death of anotherE.O. Wilson discussing his research into how ants determine when another ant is dead.(more)See all videos for this articleAnts are social insects, and the colony is a family community of which every ant is an integral unit. Apart from the community, any one individual cannot properly function or survive, and the larvae are completely dependent upon the continuous care of the adults. There are generally three castes, or classes, within a colony: queens, males, and workers. Male ants play no part in everyday nest activities. They live only for a short time, occur in limited numbers, and are virtual parasites of the colony, which must feed them. The fertile female, the queen, performs only one task: egg laying. The life cycle of the ant has four stages—egg, larva, pupa, and adult—and typically spans a period of 8 to 10 weeks for worker ants. bullet ant queenBullet ant (Paraponera clavata) queen. The presence of wings indicates that she is a virgin queen; her wings will be lost once she has mated.(more)red ant queenA red ant queen with a cluster of her eggs.(more)At certain times of the year, the winged males and virgin queens fly into the air, where the queen mates with a single male. During the flight he transfers to her seminal receptacle all the sperm she will require for the rest of her life, which may be as long as 15 years. The males die soon afterward, and the fertilized queen establishes a new nest or takes over the current nest. Her wings then drop off, and the bulky wing muscles degenerate, providing nutritive materials from the breakdown of the muscle tissue. As soon as the wings have fallen, her ovaries become functional, and egg laying begins. In primitive species, the queen leaves the nest and forages for food for the larvae. In more advanced forms, the queen rarely leaves the nest. She feeds so-called nutrition eggs or other food stores within her own body to the first brood. The larvae that survive in the nest develop into dwarf workers, which forage outside the nest for food to nourish additional larvae. honey ant repletesHoney ant repletes (Myrmecocystus).(more)The essential work in the ant society—such as building the nest, feeding and tending the brood, and defending the nest—is performed by the workers, all of whom are female. The workers can be differentiated morphologically and physiologically as soldiers, outside workers, inside workers, and nest builders. In the division of labor among some ant forms, highly specialized types of polymorphism have developed. The Cryptocercus ants, for example, make nests in hollow stems of plants, then bore a circular entrance that remains under constant surveillance by special guards whose heads are modified into pluglike structures that fit the entrance. Each guard is relieved after several hours, and another guard takes its place; entrance guards are useless for other tasks. Honey ant repletes are a special type of worker that are fed so much that the size of their abdomens is greatly increased. Unable to walk, they hang as living honey jugs from the ceiling of the nest, to be used as a food source when fresh food is scarce.
Get a Britannica Premium subscription and gain access to exclusive content.
Subscribe Now
ant hillsAnt hills in a field.(more)ant colonyBlack ants clustered around the entrance to their nest.(more)Most ants live in nests, which may be located in the ground or under a rock or built aboveground and made of twigs, sand, or gravel. Many subterranean nests are quite extensive, with a multitude of tunnels and specialized chambers. Carpenter ants (Camponotus) are large black ants common in North America that live in old logs and timbers. Some species live in trees or in the hollow stems of weeds. Tailor, or weaver, ants, found in the tropics of Africa (e.g., Tetramorium), make nests of leaves and similar materials held together with silk secreted by the larvae. Dolichoderus, a genus of ants that are found worldwide, glues together bits of animal feces for its nest. The widely distributed pharaoh ant (Monomarium pharaonis), a small yellowish insect, builds its nest either in houses, when found in cool climates, or outdoors, when it occurs in warm climates. Food The food of ants consists of both plant and animal substances. Many ants are generalists and utilize a wide range of organic substances for food. Worker ants forage daily, and collected food and water is brought to the larvae and mature ants in the nest. They frequently use scent marks, which they place on their pathways, to find their way back to the nest and direct other colony members to a food source. worker antsA swarm of ants cooperating to collectively move a dead insect on the forest floor.(more)Some ant species are hunters and scavengers. Bullet ants, for example, forage for live spiders, frogs, and insects, including grasshoppers, beetles, and katydids, or their carcasses. Certain species, including those of the genus Formica, often eat the eggs and larvae of other ants or those of their own species in other nests. Sahara desert ants (genus Cataglyphis) scavenge for dead insects on the scorching sand and salt-pan terrain of the Sahara; they can tolerate surface temperatures of 60 °C (140 °F) or higher for short periods, making them one of the most heat-tolerant groups of insects known. red imported fire antsRed imported fire ants (Solenopsis invicta).(more)Benefits and dangers of pine resin to antsWood ants collecting dried resin from a pine tree, with one ant becoming trapped in the sticky substance.(more)See all videos for this articleSome species eat the liquid secretions of plants, either directly or indirectly from the bodies of other insects. Many ants collect nectar from floral or extrafloral nectaries, and some utilize resins and saps. A number of ants, known as herder ants (Lasius niger and others), protect and carefully tend to herds of aphids, from which the ants collect honeydew (a by-product of digestion secreted by certain aphids). The honey ants (Myrmecocystus, Camponotus, and others) store honeydew in the distended abdomens of specialized workers. Some genera (Leptothorax) eat the honeydew that has fallen onto the surface of a leaf. The so-called Argentine ant (Iridomyrmex humilis) and many fire ants (Solenopsis) also eat honeydew. leafcutter antLeafcutter ant carrying a leaf fragment.(more)Follow the trails of leafcutter ants stripping rainforest foliage to cultivate fungus-based food in their nestLeafcutter ants and many other ant species are common in tropical rainforests.(more)See all videos for this articleHarvester ants (Messor, Pogonomyrmex) store grass, seeds, or berries in the nest, whereas ants of the genus Trachymyrmex of South America eat only fungi, which they cultivate in their nests. The Texas leafcutter ant (Atta texana) is a pest that often strips the leaves from plants to provide nourishment for its fungus gardens. Notable ant behaviors The social behavior of ants, along with that of honeybees, is the most complex in the insect world. The group is also extremely diverse, with any number of foraging, nesting, and social behaviors. mutualism: acacia antsMutualism between acacia ants (Pseudomyrmex ferruginea) and the bullhorn acacia (Vachellia cornigera), or swollen thorn acacia. The plant provides food and shelter to the ants, and the ants defend the plant against browsing animals.(more)Acacia ants (Pseudomyrmex ferruginea) inhabit the bullhorn acacia (or bullhorn wattle; Vachellia cornigera). The ants obtain food and shelter, and the acacia depends on the ants for protection from browsing animals, which the ants drive away. Neither member can survive successfully without the other, exemplifying obligative mutualism. Slave-making ants, of which there are many species, have a variety of methods for “enslaving” the ants of other species. The queen of Bothriomyrmex decapitans of Africa, for example, allows herself to be dragged by Tapinoma ants into their nest. She then bites off the head of the Tapinoma queen and begins laying her own eggs, which are cared for by the “enslaved” Tapinoma workers. Workers of the slave-making ant Protomognathus americanus raid nests of Temnothorax ants, stealing the latter’s pupae. The pupae are raised by P. americanus to serve as slaves, and, because the Temnothorax pupae become imprinted on the chemical odor of the slave-making ants, as adults the captive ants forage and routinely return to the slave-making ant nest. Some species live in the nests of other species as parasites. In these species the parasite larvae are given food and nourishment by the host workers. Wheeleriella santschii is a parasite in the nests of Monomorium salomonis, the most common ant of northern Africa.
Observe an Eciton army ant colony migrating by night and forming a bivouac nest entirely out of themselvesArmy ants (genus Eciton) migrating and gathering in a bivouac.(more)See all videos for this articleArmy ants, of the subfamily Dorylinae, are nomadic and notorious for the destruction of plant and animal life in their path. The army ants of tropical America (Eciton), for example, travel in columns, eating insects and other invertebrates along the way. Periodically, the colony rests for several days while the queen lays her eggs. As the colony travels, the growing larvae are carried along by the workers. Habits of the African driver ant (Dorylus) are similar. The Editors of Encyclopaedia Britannica This article was most recently revised and updated by Melissa Petruzzello.
Ants, facts and information
Skip to contentNewslettersSubscribeMenuA leafcutter ant photographed at St. Louis Zoo in Missouri. There are up to tens of thousands of ant species in the world—and 47 of them are different species of leafcutter ants, which are found in the Western Hemisphere.Please be respectful of copyright. Unauthorized use is prohibited.AnimalsReferenceAnts, facts and informationShareTweetEmailCommon Name: AntsScientific Name: FormicidaeType: InvertebratesDiet: OmnivoreGroup Name: Army, colonyAverage Life Span: Several weeks to several yearsSize: 0.08 to 1 inchesSize relative to a paper clip: What are ants?Ants are common insects, but they have some unique capabilities—including their legendary communication skills that allow their colonies to function as superorganisms.There are 20 quadrillion ants on Earth—that’s 2.5 million ants for every human. Known ant species (members of the family Formicidae) number over 12,000, and some experts estimate upwards of 20,000 exist. They can be found almost anywhere in the world, with the exception of Antarctica, Iceland, Greenland, and some island nations.(How do ants make “milk”?)Invasive ant species are becoming a problem, however, with research showing that over 500 species of ants were found in countries where they aren’t native, having hopped a ride with humans, or our cargo and goods. These alien ants can change the dynamics of an ecosystem, consume resources, and kill other organisms. This damage adds up: From 1930 to 2021, invasive ants caused an estimated $51 billion in economic losses.Texas leafcutter ants (Atta texana), feed on an apple core at Dallas Zoo.Please be respectful of copyright. Unauthorized use is prohibited.AppearanceAnts range in size from the miniscule up to one inch long, and usually appear black, brown, red, or yellow.Ants look much like termites, and the two are often confused. Ants can be identified by their elbowed antennae, and narrow "waist" between the abdomen and thorax. Some ants have wings, which are longer in the front and shorter by their hind legs. The presence of wings indicates an ant’s fertility—ants with wings are either queens or the drones whose job it is to mate with them.Diet and behaviorEnthusiastically social insects, ants typically live in structured nest communities that may be located underground, in ground-level mounds, or in trees. Carpenter ants, which include more than a thousand species in the genus Camponotus, nest in wood and can be destructive to buildings. (Much like termites, which cause far more severe damage.) Some species, such as army ants, defy the norm and do not have permanent homes.1:09WATCH: "Paramedic" Ants Save the Wounded in Termite WarsMegaponera analis, a small black ant species native to sub-Saharan Africa, wages war on termite nests. Attacking ants can have limbs ripped off or even be decapitated by the termites. Instead of leaving the hurt ants behind, other ants will carry them back home where they can heal and participate again in future raidsA single ant colony can contain hundreds of thousands of individual ants. Communities are headed by a queen or queens—some polygynous species can have as few as two or up to thousands of queens. Queens lay thousands of eggs to ensure the survival of the colony. In some species, male ants (known as drones) often have only one role—mating with the queen. They often die shortly after. Worker ants, the most visible colony members, are females that never reproduce, but instead forage for food, care for the queen's offspring, work on the nest, and protect the community. Some workers can carry 50 times their own body weight. (Why invasive ants are a bigger threat than previously thought.)Ants communicate and cooperate by secreting pheromones, or scent chemicals, that can alert others to danger or lead them to a promising food source. They typically eat nectar, seeds, fungus, or insects. However, some species have diets that are more unusual. Army ants, with their large mandibles and painful stings, may prey on reptiles, birds, or even small mammals.Please be respectful of copyright. Unauthorized use is prohibited.Please be respectful of copyright. Unauthorized use is prohibited.Left: Honeypot ant (Myrmecocystus mexicanus) is photographed with its swollen golden abdomen.Right: A red velvet ant (Dasymutilla occidentalis) at the Audobon Nature Institute in New Orleans.Ant colonies are so tightly knit and efficient, they can pass useful knowledge between generations, which some experts believe constitutes a colony “memory.” This kind of communal knowledge is essential for defense, so ants can easily differentiate friendly and hostile forces.Unusual antsWithin the many thousands of known ant species, there are many that fill unique ecological roles, resulting in the development of special physical characteristics and interesting behavior.For example, ants have a variety of biological defenses. Fire ants bite and sting with a venom called solenopsin. They can also survive floods by clumping together to float on the water’s surface. Other species, like the newly discovered Pheidole drogon, have evolved to grow spikes or spines from their exoskeletons.(How these ants help find fossils.)One Amazon species (Allomerus decemarticulatus) cooperatively builds extensive traps from plant fiber. When an insect steps on one of the trap’s many holes, hundreds of ants inside use the openings to seize it with their jaws.Another species, the yellow crazy ant (Anoplolepis gracilipes), is capable of forming so-called supercolonies that house hundreds or even thousands of queens. On Australia’s Christmas Island, the accidental introduction of yellow crazy ants in the early 20th century has led to a destructive infestation. The ants are a significant threat to the island’s endemic population of red crabs, displacing the crustaceans from their burrows or even killing them as they pass by ant nests during their annual migration from the forest to the coast.Did you know?Ants have a symbiotic relationship with a group of plants called Myrmecophytes, which provide food and the ants protect these plants from pests.— ScienceDirectThere are several wasp species that closely resemble ants, including the velvet ant, which is actually a hairy, wingless wasp.— University of Nebraska-LincolnAncient ants found perfectly preserved in amber have given us invaluable clues to their evolution. One specimen of an ant caught hunting is believed to be 99 million years old. — BBC Science FocusShareTweetEmailGo FurtherAnimalsRare gray whale spotted in the Atlantic—and it's only the beginningAnimalsRare gray whale spotted in the Atlantic—and it's only the beginningWhy 'funga' is just as important as flora and faunaScienceWhy 'funga' is just as important as flora and faunaTermite fossils prove mating hasn't changed in 38 million yearsAnimalsTermite fossils prove mating hasn't changed in 38 million yearsPlay to find out which birds dominate at your feeder—and whyAnimalsPlay to find out which birds dominate at your feeder—and whyHow do fireflies get their glow? We finally have some answers.AnimalsHow do fireflies get their glow? We finally have some answers.Bird flu is spreading from pole to pole. Here’s why it matters.AnimalsBird flu is spreading from pole to pole. Here’s why it matters.EnvironmentMushroom leather? The future of fashion is closer than you think.EnvironmentMushroom leather? The future of fashion is closer than you think.This deadly fungus is hitchhiking its way across the worldEnvironmentThis deadly fungus is hitchhiking its way across the worldWhy 'funga' is just as important as flora and faunaScienceWhy 'funga' is just as important as flora and faunaThis exploding mine holds a treasure that may change the worldEnvironmentThis exploding mine holds a treasure that may change the worldHow animals are adapting to the rise of wildfiresEnvironmentHow animals are adapting to the rise of wildfiresFast fashion goes to die in this Chilean desertEnvironmentFast fashion goes to die in this Chilean desertHistory & CultureWho was the real Calamity Jane?History & CultureWho was the real Calamity Jane?See how these fierce dinos evolved in our pages over 100 yearsScienceSee how these fierce dinos evolved in our pages over 100 yearsHow did this female pharaoh survive being erased from history?History MagazineHow did this female pharaoh survive being erased from history?How Black artists helped make country music what it is todayHistory & CultureRace in AmericaHow Black artists helped make country music what it is todayThese 3 samurai women were heroes of shogun era JapanHistory & CultureThese 3 samurai women were heroes of shogun era JapanLincoln was killed before their eyes—then their own horror beganHistory & CultureLincoln was killed before their eyes—then their own horror beganScience'Exploding head syndrome'? Inside a mysterious sleep conditionScience'Exploding head syndrome'? Inside a mysterious sleep conditionWhat lifting weights does to your body—and your mindScienceMind, Body, WonderWhat lifting weights does to your body—and your mindMushroom leather? The future of fashion is closer than you think.EnvironmentMushroom leather? The future of fashion is closer than you think.Fungi could be the key to major cancer research breakthroughsScienceFungi could be the key to major cancer research breakthroughsWhy 'funga' is just as important as flora and faunaScienceWhy 'funga' is just as important as flora and faunaWhat is a lunar eclipse—and when will the next one happen?ScienceWhat is a lunar eclipse—and when will the next one happen?TravelKingston is making a comeback as culture capital of JamaicaTravelKingston is making a comeback as culture capital of JamaicaYou can watch wild horses roam free in this UK national parkTravelYou can watch wild horses roam free in this UK national parkWhere to stay in Zanzibar, Tanzania's 'spice island'TravelWhere to stay in Zanzibar, Tanzania's 'spice island'To see the real Los Angeles, visit its historic movie theatersTravelTo see the real Los Angeles, visit its historic movie theatersThis comedian went on a quest to find the end of the worldTravelThis comedian went on a quest to find the end of the worldHistory is alive on these European islandsTravelHistory is alive on these European islandsLegalTerms of UsePrivacy PolicyYour US State Privacy RightsChildren's Online Privacy PolicyInterest-Based AdsAbout Nielsen MeasurementDo Not Sell or Share My Personal InformationOur SitesNat Geo HomeAttend a Live EventBook a TripBuy MapsInspire Your KidsShop Nat GeoVisit the D.C. MuseumWatch TVLearn About Our ImpactSupport Our MissionMastheadPress RoomAdvertise With UsJoin UsSubscribeCustomer ServiceRenew SubscriptionManage Your SubscriptionWork at Nat GeoSign Up for Our NewslettersContribute to Protect the PlanetFollow usNational Geographic InstagramNational Geographic FacebookNational Geographic TwitterNational Geographic YoutubeNational Geographic LinkedinNational Geographic TiktokNational Geographic RedditUnited States (Change)Copyright © 1996-2015 National Geographic SocietyCopyright © 2015-2024 National Geographic Partners, LLC. All rights reserved
Types of Ants With Identification and Pictures (Identification Chart)
Home
About Us
Contact Us
Privacy Policy
Disclaimer
Flowers
Houseplants
Produce
Trees
Gardening And Landscaping
Succulents
Shrubs
Types of Ants With Identification and Pictures (Identification Chart)
By Jessica Nolan, Gardening Expert
Animals
Share on Email
Share on Pinterest
Share on Facebook
Share on Twitter
Share on LinkedInAnts are a type of small insect with three distinct body parts, six legs, and two antennae. Black ants are the most common type; however, ant species can also be dark brown, tan, or red colors. Some types of swarming ants also develop wings. But most ants are tiny creepy crawlies that get around by walking.
Small ants and large ants are social insects that live in large colonies. Therefore, if you spot a few ants in your home or garden, the chances are that there are hundreds of ants nearby. Ants can become a nuisance pest in homes or gardens. These tiny bugs can inflict painful stings and chew through wood, electrical cables, or insulation.
This article is a comprehensive guide to identifying common types of ants you’re likely to encounter at home or in the yard. Knowing how to identify an ant species can help you understand how to get rid of the tiny crawling insects.
How to Identify Ants
Ant identification is by their color, size, and body shape. Ants have three pairs of legs and three identifiable body parts — the head, thorax, and abdomen. Some ants have large, bulbous heads and abdomens with a thin thorax. The common names of some ants describe their appearance and habits.
For example, small odorous ants, black household ants, little black ants, crazy ants, and white-footed ants are types of ants where their common names help identify their characteristics.
Types of Ants With Pictures and Names — Identification Guide
Let’s look in detail at the identifying features of common types of ants.
Pharaoh Ant (Monomorium pharaonis)
A close up picture of the miniature tiny Pharaoh ant
The pharaoh ant is a small light brown or orange-yellow ant that can become a significant pest in buildings. The tropical ant measures around 0.078” (2 mm) long. The tiny brown ant has a large dark brown to black bulbous abdomen and two distinctive club antennae.
Pharaoh ants live in large colonies and have several red-colored queen ants per colony. The ants feed on sugary and greasy foods and can infest wall voids, furniture, and under floors.
It is said that an ant bait consisting of 1 percent boric acid and sugary water can kill pharaoh ants.
Ant Identification
Pharaoh ants have slender, light brown, almost translucent bodies with a large ball-like blackish abdomen.
Small Odorous House Ant (Tapinoma sessile)
Enlarged picture of the small odorous house ant that emits foul smell when crushed
The small odorous house ant is black or brown and measures between 0.06” and 0.12” (1.5 – 3.2 mm). The tiny ants are recognized by their abdomen (gaster) that is closely joined to the thorax. This feature makes the abdomen appear pointed downward.
Odorous house ants typically feed on honeydew — a common sign of aphids on houseplants or garden shrubs. The best way to prevent odorous ants from infesting homes is to keep vegetation and plants growing away from buildings.
The small odorous house ants got their common name because of the nasty smell they emit when crushed. These small black ants are also called the stink ant for the same reason.
Ant Identification
The brown odorous house ant is identified by its oval abdomen, triangular head, and segmented antennae.
Black Household Ant (Ochetellus glaber)
The small black household ant can also be dark brown, depending on the species
The tiny black household ant can be jet black or dark brown, depending on the subspecies. Black house ants measure 0.08” to 0.11” (2 – 3 mm) long. The ants have curved antennae, and their legs may appear yellow. Up close, pictures of the black ants show they have a wrinkled face and large compound eyes.
The tiny black or brown household ants feed on sweet liquids and you’ll often find these common household invaders in the bathroom, kitchen, or laundry room. Outdoors, the ants feed on honeydew, nectar, and other insects. The black ants are common in Florida and other states in the southern US.
Ant Identification
The black household ant is a tiny ant that can be brownish-black or dark brown in color with a narrow waist, slender thorax, and large abdomen.
Little Black Ant (Monomorium minimum)
The little black ants are common in dark places in the house and garden
The little black ant is a type of little jet-black ant measuring 0.04” to 0.078” (1 – 2 mm) long. The tiny black ant is recognized by its two-segmented waist (pedicel), clubbed antennae, stinger, and ball-like, shiny black abdomen. The tiny black ants live in medium-sized colonies.
Little black ants are found throughout the United States, especially in California. The small black ants inhabit dark places in yards under logs and piles of bricks. You’ll find the little black ants inside homes in wall voids, behind baseboards, or in masonry crevices.
Ant Identification
Little black ants are identified by their shiny black bodies, club antennae, and tiny size.
Black Carpenter Ant (Camponotus pennsylvanicus)
The big black carpenter ant can damage wooden furniture and structures
Carpenter ants are some of the largest black ants, measuring 0.23” to 0.5” (6 – 13 mm) long. The large black ants have whitish hairs on their bodies and a striped abdomen. Additionally, the ants have characteristic elbowed antennae and six brownish-black legs attached to the thorax.
Carpenter ants get their name from their habit of excavating dead wood. Unfortunately, this means that the destructive black bugs can damage furniture and even cause structural damage over time. To prevent a carpenter ant invasion, reduce humidity and keep moist wood away from structures.
Ant Identification
The black carpenter ant is a large ant with a dull black body, fuzzy abdomen, and smooth thorax.
Pavement Ant (Tetramorium)
Pavement ants include many species. In the picture: Tetramorium caespitum
The pavement ant is a common dark brown ant that can sting humans. The slender brown ant has a rounded head, oval brown abdomen, and stinger. These small brown insects measure 0.1” (3 mm) long and are difficult to spot due to their minuscule size.
Pavement ants thrive in sand and soil and prefer to live under sidewalks, driveways, and patios. Inside homes, you may find pavement ant nests in basements. Unfortunately, the ants tend to nest underground, making getting rid of the brown bugs challenging.
Ant Identification
A pavement ant is recognized by its dark brown to black body with recognizable ridges and fine hairs on its head and thorax. Pictures of the brown ant show it has a stinger at its tail end.
Black Garden Ant (Lasius niger)
The small black garden ant is common in many parts of the world
Called the common black ant, this small black insect measures around 0.2” (5 mm) long. Sometimes a dark brown color, the common garden ant has identifiable erect hairs on its antennae and top leg sections. In addition, some black ants have a faint stripe around their abdomens and a thin waist.
The black garden ant is the species you are likely to find in your yard. The male flying ants can sometimes find their way into homes. This can attract hordes of more ants to come foraging for sweet foods.
Ant Identification
The common black garden ant has a recognizable black, bulbous abdomen attached to the thorax by a thin waist and six dark brown legs.
Related reading: How to Control Ants in the Garden
Banded Sugar Ant (Camponotus consobrinus)
The orange-brown and black banded sugar ant is relatively large and is found in forests and houses
The banded sugar ant is dark brown to black and has a light-colored rusty brown band around its abdomen. Measuring 0.2” to 0.6” (4 – 16 mm), the large brown ant has a shiny black triangular head, egg-shaped brown and black abdomen, and coppery-brown legs.
The banded sugar ant is native to Australia. You will also find colonies of sugar ants nesting under shrubs, dead wood, and in the soil. The black and orange-brown ants also construct nest mounds above ground.
Ant Identification
A banded sugar ant appears to have a rusty brown band on its black abdomen. The orange-brown ant has a black head and brown legs.
Yellow Meadow Ant (Lasius flavus)
The little yellow meadow ant feeds on aphids honeydew
The yellow meadow ant is a common small yellowish-brown ant species, measuring 0.78” to 0.15” (2 – 4 mm) in length. The meadow ant has a yellow head, thorax, and abdomen (gaster), which is darker at the edges. The antennae are 12-segmented and end in a three-segmented club.
The yellow meadow ants are not particularly aggressive and often nest under rocks or slabs. These tiny yellowish-brown ants also protect aphids as they like to feed on the honeydew. You may find them in gardens when they create a soil dome for a nest.
Ant Identification
The yellow meadow ant has an identifiable yellowy-brown color and an egg-shaped yellowish abdomen with fine setae (hairs) at its tail end.
Fire Ant (Solenopsis spp.)
The fire ant is a small red ant that can cause a painful sting
The fire ant is a common stinging slender ant identified by its dark brown to red color and aggressive behavior. The nasty small red ants have a copper-brown head and thorax and dark red abdomen. Additionally, they have two ten-segmented antennae with two-segmented clubs. The small red ants measure 0.8” to 0.23” (2 – 6 mm).
Fire ants can be found in Texas, Alabama, and other states in the southern US. The tiny fire ants are known for their painful stings that cause pain, like being burned by fire. Red fire ants nest in the soil or dome-shaped mounds and can attack when disturbed.
Ant Identification
The small red fire ant is easily identified by its reddish-brown color, two black eyes, exceptionally slender thorax, and large reddish-brown abdomen.
Crazy Ant (Paratrechina)
A magnified photo of the long legged crazy ant (Paratrechina longicornis)
The crazy ant is a species of small dark brown to black ant with whitish setae and extremely long antennae. These tiny, fast-moving ants measure 0.9” to 0.11” (2.3 – 3 mm) and have an unevenly rounded thorax. Small crazy ants are known for their erratic, speedy movements.
Crazy ants are a species of tropical ant commonly found outdoors in Florida and the Gulf states. However, the reddish-brown crazy ants also live in the house in temperate climates. As a result, they are a serious nuisance pest and forage on household foods, fruit, plant honeydew, and seeds in many areas.
These crazy ants can be extremely difficult to get rid of.
Ant Identification
A crazy ant is distinguished from other ants by a long pair of antennae, brownish-black color, and thin legs with relatively few hairs on them.
Ghost Ant (Tapinoma melanocephalum)
The tiny ghost ant is identified by its dark brown head and beige translucent abdomen
The ghost ant is a species of tiny dark brown and creamy white ant common in Florida. The ghost ant is identified by its dark brown head and thorax, pale translucent abdomen with faint dark patterns. These difficult-to-spot tiny ants measure 0.05” to 0.08” (1.3 – 2 mm) long.
Ghost ants are a common household pest in Florida, Texas, and throughout the US. Ghost ants are common household invaders and they can be found in kitchens, bathrooms, and other humid areas. They typically nest outdoors but will forage indoors if conditions are suitable and moisture is present.
Ant Identification
A ghost ant is identified as a small brown ant with a yellowish abdomen, milky white antennae without clubs, and six transparent legs.
Red Wood Ant (Formica rufa)
The red wood ant has a black abdomen and creates large nests in the forests
The red wood ant is identified by a black abdomen and red thorax, black and red head, and long black legs. This type of small black and red ant species measures 0.17” to 0.35” (4.5 – 9 mm). The red ants have short, elbowed antennae.
Also called thatching ants, wood ants, mound ants, and field ants, the ants are found in deciduous forests throughout the United States. The small ants create large ant hills, sometimes several feet high. The busy ants typically feed on aphids and are important in forestry management.
Ant Identification
The small red wood ant is identified by a reddish-brown head and thorax, a dark brown or black abdomen, and black legs. The red ants are easy to spot in forests because they appear in large numbers.
Argentine Ant (Linepithema humile)
The brown Argentine ant can be a serious household pest
The Argentine ant is a tiny, brownish-colored ant measuring 0.06” to 0.11” (1.6 – 2.8 mm) long. The light to dark brown ant has a distinguishable single node between the abdomen and thorax and two bands over the abdomen. Another feature of the Argentine ant is its tan legs and segmented antennae.
Argentine ants are an invasive species and a significant pest in urban areas. The pesky insects invade homes during dry or hot weather and forage for food in kitchens. There can sometimes be hundreds of these brown ants scurrying over floors.
Ant Identification
Argentine ants are small, light to dark brown ants with pale brown legs and club-less antennae with 12 segments.
Leafcutter Ants
Leafcutter ants can be found in South and Central America, and southern USA
Leafcutter ants comprise of over 40 ant species that cut leaves with their mandibles (mouthparts) to carry them back to the nest. The common identifying feature of leaf-chewing ants is how hundreds of the insects carry leaf parts in line, one after the other.
In some places, leafcutter ants can be serious agricultural pests. For example, some species can strip citrus trees bare in less than 24 hours.
The species Atta sexdens, is a type of red ant with a broad, rounded head, slender thorax, and small, round abdomen. The red ant measures around 0.04” to 0.08” (1 – 2 mm) long. The largest of the red leafcutter ants can have a head over 0.11” (3 mm) wide.
Red Harvester Ant (Pogonomyrmex barbatus)
The red harvester ant has copper-brown body and are often confused with fire ants
The red harvester ant has a large, red to brownish body with a rounded, bulbous abdomen and elongated thorax. These copper-brown large ants measure 0.19” to 0.27” (5 – 7 mm) long. Identifying features of the harvester ants are fine setae on its back and abdomen, small black eyes, and red elbowed antennae.
Because of their red appearance, red harvester ants are easy to mistake for unrelated fire ants. Species of harvester ants are common in Texas and other states throughout the US and Canada. Although these are red stinging ants, they rarely sting.
Ant Identification
The red harvester ant is a large reddish-brown ant with two 12-segmented antennae and fine hairs on its head. The red ants are easy to spot in a landscape because they construct large, mounded nests.
White-Footed Ant (Technomyrmex albipes)
The small white-footed ant can be found in forests as well as homes
The white-footed ant is a small black ant with six black legs and creamy-white tips. Typically, the dull black ants measure 0.08” to 0.16” (2 to 4 mm) long. Identifying features of the white-footed ants are two 12-segmented antennae, five abdominal segments, and fine setae on its back.
White-footed ants look like Argentine ants and are often confused with crazy ants. The black and white ant is common throughout Florida, Louisiana, Georgia, and South Carolina. It is considered a pest when it infests indoor spaces.
Ant Identification
The white-footed ant has identifiable white legs and a black head, thin thorax, and egg-shaped brownish-black abdomen.
Pyramid Ant (Dorymyrmex bureni)
The pyramid ant has orange-reddish body and emit bad odor when crushed
The pyramid ant is medium-sized with a pale orangey-red or pale-yellow body, club-less antennae, no sting, and a slender body. The ant is not considered aggressive. However, it emits a foul stench of rotten coconuts when crushed. The small to medium-size ants measure 0.8” – 0.16” (2 – 4 mm) long.
Common to Florida, the pyramid ant is not classified as an aggressive ant, unlike red fire ants. Instead, the small reddish-brown ants with their tiny black eyes feed on honeydew, other insects, and live outdoors in nests they construct in mounds of sand.
Ant Identification
The pyramid ant is a slender orange-red ant with six thin orange legs, long 12-segmented antennae without a defined club, and a pyramid-shaped projection on the thorax.
Learn How to Control Ants in the Garden.
Related articles:
Types of Insects with Pictures and Names
Types of Small Black Bugs – Identification
Read NextTypes of Gray Bugs: Silverfish, Stink Bugs and More (With Pictures) - Identification GuideSmall Brown Beetles (In the House and Outside) - With Pictures and IdentificationCommon House Spiders (With Pictures) - Identification GuideTypes of Beautiful Moths (With Pictures) - Identification GuideTypes of Ants With Identification and Pictures (Identification Chart) was last modified: March 9th, 2022 by Jessica Nolan, Gardening Expert
Share on Email
Share on Pinterest
Share on Facebook
Share on Twitter
Share on LinkedIn
© 2024 Leafy Place
Powered by WordPress
FlowersHouseplantsProduceTreesGardening And LandscapingSucculentsShrubs
× Close Panel
Ant - Description, Habitat, Image, Diet, and Interesting Facts
HomeAnimal IndexComplete List of AnimalsA – GAnimals that start with AAnimals that start with BAnimals that start with CAnimals that start with DAnimals that start with EAnimals that start with FAnimals that start with GH – NAnimals that start with HAnimals that start with IAnimals that start with JAnimals that start with KAnimals that start with LAnimals that start with MAnimals that start with NO – TAnimals that start with OAnimals that start with PAnimals that start with QAnimals that start with RAnimals that start with SAnimals that start with TU – ZAnimals that start with UAnimals that start with VAnimals that start with WAnimals that start with XAnimals that start with YAnimals that start with ZDogsParks and ZoosCitationContact Us Search HomeAnimal IndexComplete List of AnimalsA – GAnimals that start with AAnimals that start with BAnimals that start with CAnimals that start with DAnimals that start with EAnimals that start with FAnimals that start with GH – NAnimals that start with HAnimals that start with IAnimals that start with JAnimals that start with KAnimals that start with LAnimals that start with MAnimals that start with NO – TAnimals that start with OAnimals that start with PAnimals that start with QAnimals that start with RAnimals that start with SAnimals that start with TU – ZAnimals that start with UAnimals that start with VAnimals that start with WAnimals that start with XAnimals that start with YAnimals that start with ZDogsParks and ZoosCitationContact Us Animals Network HomeAnimal IndexComplete List of AnimalsA – GAnimals that start with AAnimals that start with BAnimals that start with CAnimals that start with DAnimals that start with EAnimals that start with FAnimals that start with GH – NAnimals that start with HAnimals that start with IAnimals that start with JAnimals that start with KAnimals that start with LAnimals that start with MAnimals that start with NO – TAnimals that start with OAnimals that start with PAnimals that start with QAnimals that start with RAnimals that start with SAnimals that start with TU – ZAnimals that start with UAnimals that start with VAnimals that start with WAnimals that start with XAnimals that start with YAnimals that start with ZDogsParks and ZoosCitationContact Us ArthropodaHymenopteraFormicidaeInsectaEuarthropodaAntBy Animals Network TeamThis impressively diverse group of animals has developed hundreds of different adaptations to survive in many different habitats. Scientists estimate that we have discovered only slightly more than half of the ant species in the world. The species that we have discovered are simply fascinating. Read on to learn about the ant. Description of the AntAll ants have elbowed antenna, which are bent like elbows. Their bodies consist of a head, alitrunk, petiole, and gaster. The alitrunk is the first segment of the body, which is attached to the head. The alitrunk is followed by a characteristically narrow segment, called the petiole. Finally, the gaster is the large hind segment of the ant, and is typically ended with a stinger.Interesting Facts About the Ant With over 12,500 identified species, and an estimated 22,000 species worldwide, there are virtually endless facts about ants. Different species have different traits that make them unique. Some of the most unique traits are found in only small groups or individual species.Fungal Farms – There is an entire group of ants, consisting of 47 species, that survives by farming its own food. The leafcutter ants trim segments of fresh vegetation, and carry it to their nests. They use this vegetation to grow their own fungus farms! The fungus is fed to the ant larvae, and the ants feed the fungus and keep it free from bacteria and microbes. This coexistence is called a symbiotic relationship, where both creatures benefit.Guard Ants – In another display of symbiosis, acacia ants and acacia trees work together to survive. The ants use the hollow thorns of the tree to nest in, and feed on the sap of the tree. In return for the free home and food, the acacia ants protect the tree from any potential hazards, such as chewing insects or hungry herbivores. Nothing like a swarm of angry ants to drive you to feed on a different tree!Ants as Bodyguards – Speaking of providing protective details, another group of ants partners with aphids in another symbiotic relationship! These ants follow and protect the aphids as they forage for sap, and in return the aphids produce a highly nutritious nectar secretion for the ants. To release this secretion, the ants use their antennae to stroke the backs of the aphids.Caterpillar Caper – Just like the relationship between ants and aphids, a different species of ants receives preferential treatment from the lycaenid caterpillar. In a similar display, the ants are fed nectar from the caterpillar’s secretions, and in return the ants protect the caterpillar. They will even swarm onto the caterpillar’s back to discourage birds from preying on their caterpillar, as ants taste unpleasant to birds.Habitat of the AntAnts can be found in virtually any ecological niche, and live in a wide range of habitats. They can be found in urban settings, tropical rainforests, deciduous forests, deserts, and many, many more.Distribution of the AntJust like their wide habitat distribution, ants can be found across many different locations as well. They naturally occur in virtually every landmass on earth. The only places without native species of ants are Antarctica, Polynesia, Iceland, Greenland, and Hawaii. In some of these locations invasive ant species have been introduced.Diet of the AntMost ant species are omnivores, and will feed on both plant and animal matter. Many ants feed on the nectar of plants, other insects (including those many times their own weight), fruits and vegetables, human garbage, and meat from carcasses of animals.Ant and Human InteractionAnt and human interaction is generally quite negative. When ants and humans cross paths, bites and stings frequently occur. There are a number of invasive ant species, which pose a problem to native flora and fauna. Humans also pose a danger to native ant species, through direct extermination or indirect activity like land development and pollution.DomesticationWe have not domesticated ants, but ants have domesticated aphids!Does the Ant Make a Good PetAnt farms are a frequent children’s plaything, and can be a great learning activity. Ants as pets are a hands-off animal, and they should not be handled.Ant CareMany types of ants are easy to care for, and can make good pets. Ants should be provided with a substrate habitat for digging a nest, and suitable food to eat. This will range by species, but feeding is generally easiest when insects are provided rather than other food sources.Behavior of the AntMany different species of ants have very complex social structures. Ants identify one another using smell, and ants that are not members of the colony will be attacked. Within the ant colony there will frequently be “classes” of workers, each of which has a different job.Worker ants will have strong mandibles (jaws) and large bodies to move food and protect the nest. Drones are sexually reproducing males, who mate with queens. Queens are the only females capable of reproducing, and they do so on a mass scale.Reproduction of the AntAnts have a wide variety of reproductive strategies. In some species, male drones enter a nest and release a mating pheromone so they are not attacked. These drones will breed with the queen and the queen will produce eggs. In other species, young queens hatch and perform a mating flight with the drones. Breeding occurs in midair, and the now-pregnant queens will dig a nest and begin their own anthill.TAGSantant anatomydronesqueenred antworker antCydni Oldham RELATED ARTICLESMORE FROM AUTHOR ArachnidaBrown Recluse ArachnidaGarden Spider ArachnidaSpider Expert Recommendations Dog CareBest Dog Food Expert RecommendationBest Cat Litter Boxes Expert RecommendationBest Self Cleaning Litter Box Dog CareBest Orthopedic Dog Bed Dog CareBest Dog Food for Small Dogs Dog CareBest Dog Toys Dog CareBest No Pull Dog Harness Dog CareBest Wet Dog Food Dog CareBest Dog Food for Sensitive Stomach Dog CareBest Senior Dog FoodAnimals.NET aim to promote interest in nature and animals among children, as well as raise their awareness in conservation and environmental protection. All photos used are royalty-free, and credits are included in the Alt tag of each image.EVEN MORE NEWSRed AngusPaint HorseHouse SpiderPOPULAR CATEGORYChordata694Mammalia247Aves197Dog Breeds184Actinopterygii121Reptilia87Carnivora72Privacy PolicyContact UsAbout UsTerms and Conditions © Animals.NET 2018
x
Ants: Fun Facts About Ants & Ant Information for Kids
Glossary
Pests A to Z
>
Pest Info
Pest Guides
Pest Articles
Glossary
>
For Kids
Games
Crafts
Pest Quest Episodes
Bug Battles Episodes
"Will They Eat It?" Episodes
>
For Parents
eBooks
Downloadable Activities
Science Fair Kits
Report Writing
Pest Professor Lessons
Buggy Break
>
For Teachers
School Lesson Plans
Elementary Grade Activities
Middle School Grade Activities
Pest Quizzes
Glossary
Pests A to Z
>
Pest Info
Pest Guides
Pest Articles
Glossary
>
For Kids
Games
Crafts
Pest Quest Episodes
Bug Battles Episodes
"Will They Eat It?" Episodes
>
For Parents
eBooks
Downloadable Activities
Science Fair Kits
Report Writing
Pest Professor Lessons
Buggy Break
>
For Teachers
School Lesson Plans
Elementary Grade Activities
Middle School Grade Activities
Pest Quizzes
Ants
Did you know? An ant can lift 20 times its own body weight.
Explore Ants
Argentine Ants
Carpenter Ants
Odorous House Ants
Pavement Ants
Red Imported Fire Ants
Ant Facts for Kids
We’re all familiar with ants, right? Those tiny black insects that steal food off picnic blankets? But did you know ants can also be big, be colors other than black, and even have wings and fly? View some interesting facts about ants for kids below, help Archibald the ant complete his mission by playing the game Archibald’s Adventure, or learn how to make an egg carton ant!
There are more than 12,000 different types of ants all over the world!
Ants can lift 20 times their own body weight. If a second grader was as strong as an ant, they could pick up a car!
Ant colonies are run by a single queen ant, who control all the other ants.
Some queen ants can live for many years and have millions of babies.
Ants don’t have ears. Ants "hear" by feeling rumbles in the ground through their feet.
Ants don’t have lungs. Air enters and leaves through tiny holes all over their body.
When ants fight, it is usually to the death!
Ants leave invisible breadcrumbs (called a pheromone trail) everywhere they go, so they know where they’ve been.
Worker ants can’t have babies. That means when a queen dies, the colony usually dies a few months later.
Ants eat different kinds of foods depending on the type of ant, including sweet sugary foods or fatty oily foods.
Ants communicate through scent, touch, motion, sound, and trophallaxis.
You can make ants on a log with healthy snacks like celery and peanut butter!
Get to know ants:
There are 3 kinds of ants in a colony: The queen, the female workers, and males.
The queen and the males have wings, the workers do not. The queen is the only ant that can lay eggs. A male ant’s job is to mate with future queen ants. Once the queen is old enough to lay eggs, she spends the whole rest of her life laying eggs!
Some worker ants act as soldiers. They protect the colony, gather food, and fight other colonies for more food and space. If they beat another ant colony, they take the other colony’s eggs. When those eggs hatch, the new ants are forced to work for the new colony. They take care of the eggs and babies, find food for the colony, and build the anthills or mounds.
Download
Pest ID Card
Argentine Ant Facts for Kids
This type of ant is from Argentina and Brazil and was probably introduced to the United States in freight ships around the 1890’s. These ants can be found in southern states as well as California, Illinois, Maryland, Missouri, Oregon and Washington.
Size: 1/16" to 1/4"
Shape: Segmented, Oval
Color: Dark brown to black and shiny
Legs: 6
Wings: Varies
Antenna: Yes
Common Name: Argentine ant
Kingdom: Animalia
Phylum: Arthropoda
Class: Insects
Order: Hymenoptera
Family: Formicidae
Species: Linepithema humile
What Do Argentine Ants Eat?:
Argentine ants like sweet meals the best but will eat almost anything. That includes meats, eggs, oils and fats. They leave a scent trail everywhere they go so they don’t waste time visiting the same place twice. Unlike most ant types, Argentine queens help with finding food.
Where Do Argentine Ants Live?: Argentine ant live in wet environments typically near a good food source. Their colonies can get huge, sometimes covering entire garden or even a whole back yard.
What Problems Do Argentine Ants Cause?: You don’t really have to worry about Argentine ants trying to hurt you. But they can contaminate food by leaving their bodily waste behind. Yuck!
What Should I Do If I Find Argentine Ants?: Talk to a grown-up right away. Since colonies can take over whole yards, it’s best to have a pest control professional deal with the situation immediately. You can find information on getting rid of Argentine ants at the official National Pest Management Association website.
Download
Pest ID Card
Carpenter Ant Info for Kids & Adults
Carpenter ants get their name because they build their nests in wood. Unlike human carpenters who help fix your house, carpenter ants can cause major damage to a house. They put huge holes in the wood of your house to build their nests. The queen of a colony starts her nest deep within wood, then raises the baby carpenter ants there, feeding them saliva and never leaving the nest or feeding herself during this time.
When baby carpenter ants grow up, they become workers with the job of gathering food to feed the next generation. The colony population grows very quickly. It can eventually produce 2,000 or more workers.
There are many types of carpenter ants throughout the U.S., measuring in size from one-quarter inch (about the width of a pencil) for a worker carpenter ant to three-quarters of an inch (about the size of a quarter) for a queen carpenter ant. Read on for more fun carpenter ant facts for kids.
Size: 5/8"
Shape: Oval
Color: Range in color from red to black
Legs: 6
Wings: Varies
Antenna: Yes
Common Name: Carpenter ant
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidea
Species: Camponotus
What Do Carpenter Ants Eat?: You may think carpenter ants eat wood, since they dig big holes into the wood. But no! They don’t eat any of it. They dig holes in wood but leave piles of it outside the entrance to their home. Instead, carpenter ants eat living and dead insects, meat, and fats and sugary foods of all kinds such as honeydew and nectar from plants.
Where Do Carpenter Ants Live?: We know carpenter ants build nests in wood, but where? What type of wood do they prefer? Carpenter ants live anywhere they can find water and moldy or damp wood, such as tree stumps, firewood or in the plants around your house.
Carpenter ants also build nests inside, usually entering buildings through wet, damaged wood, although it isn’t uncommon for them to adapt to drier environments. You know you have a colony of carpenter ants if you see piles of sawdust on the floor.
What Problems Do Carpenter Ants Cause?: The good news is that carpenter ants don’t carry disease and can’t really harm you physically. But when building a nest inside a home, they dig smooth tunnels through the wood. These tunnels weaken the wood and potentially damage the wood that keeps the house standing. This kind of damage can be very expensive to fix.
What Should I Do If I See Carpenter Ants?: If you find sawdust on the floor near walls, show a grown-up immediately. A pest control professional will need to be called to assess the situation. You can find information on getting rid of carpenter ants at the official National Pest Management Association website.
Download
Pest ID Card
Odorous House Ant Facts for Kids
Something that is odorous smells bad. The odorous house ant gets its name because it commonly nests in or around houses, and because it stinks when crushed. Imagine the smell of a rotten coconut. This is what you’ll smell if you step on one of these ants. They are from the United States, and they are very social. Colonies can include up to 100,000 ants!
Size: 1/16" to 1/8"
Shape: Segmented, oval
Color: Brown or black
Legs: 6
Wings: Varies
Antenna: Yes
Common Name: Odorous house ant
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Species: Tapinoma sessile
What Do Odorous House Ants Eat?: Odorous house ants like to eat dead insects and sugary sweets, especially melon.
Where Do Odorous House Ants Live?: Odorous house ants live in the walls and floor cracks of homes. They may also live in exposed soil and under stones, logs, mulch, debris and other items. They typically have a lifespan of several years.
What Problems Do Odorous House Ants Cause?: Odorous house ants do not pose a health threat, they can’t really hurt you. But the waste they leave behind when stealing food can contaminate that food. They can also become a huge annoyance, with colonies so large in or near your home, crawling all over your kitchen, and of course stinking up the place if you try to crush them under your shoe or an object.
What Should I Do If I Find Odorous House Ants?: Don’t squish them! There may be too many to deal with anyway. The better thing to do is have a grown-up speak with a pest control professional to determine the best way to get odorous house ants out of your house. You can find more information on finding the right pest control professional at the official National Pest Management Association website.
Download
Pest ID Card
Pavement Ant Facts for Kids
Pavement ants are so named because they like to make their nests in or under cracks in pavement. But don’t be fooled — they can also live inside buildings. They are typically found in the eastern half of the United States, California and Washington. Pavement ant colonies average 3,000 to 4,000 members and have several queens.
Size: 1/8"
Shape: Segmented, oval
Color: Dark brown to black
Legs: 6
Wings: Varies
Antenna: Yes
Common Name: Pavement ant
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Species: Tetramorium caespitum
What Do Pavement Ants Eat?: Pavement ants will eat almost anything! This includes insects, seeds, honeydew, honey, bread, meats, nuts, cheese, and even grease.
Where Do Pavement Ants Live?: Pavement ants most commonly live in soil next to and beneath pavement slabs, sidewalks, patios, and driveways. They may also build nests under a building’s foundation and hollow foundation walls.
Are Pavement Ants Harmful?: Don’t worry about pavements ants too much. They don’t pose a health threat and can’t cause too much damage to your home. They can contaminate your food, though! If you find pavement ants, make sure you get professional help to remove them.
What Should I Do If I See Pavement Ants?: As with any ants, if you see a number of pavement ants around your food or throughout your house, show a grown-up, so they can call a pest control professional. An expert will come in to review the situation and figure out the best way to solve it. This goes for if you find ants in your driveway or sidewalk as well. You can learn more about this at the official National Pest Management Association website.
Download
Pest ID Card
Red Imported Fire Ant Facts for Kids
Red imported fire ants may have a cool name, and an even cooler nickname (RIFA). But be warned: they are more aggressive than other ant species on the planet. And they have a really painful sting. They should always be avoided!
Red imported fire ants can adapt to many climates and conditions in and around their environment. For example, if the colony senses increased water levels in their nests, they will come together and form a huge ball or raft that is able to float on the water! So yes, they are cool. But very dangerous. Read on to learn more red imported fire ant information for kids.
Size: 1/8" to 3/8"
Shape: Segmented, oval
Color: Dark reddish brown
Legs: 6
Wings: Varies
Antenna: Yes
Common Name: Red imported fire ant
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Species: Solenopsis invicta
What Do Red Imported Fire Ants Eat?: Red imported fire ants primarily eat vegetation. That’s right — even though they will bite people, they can be considered vegetarians. They aren’t biting you to eat or to be mean, but to defend themselves. After all, you’re a lot bigger than they are.
Where Do Red Imported Fire Ants Live?: Red imported fire ant live in very noticeable mounds. These mounds are huge, sometimes a whole foot tall, and typically in the shape of a dome. They like to build these mounds outdoors out of soil, usually in landscape areas like grass or near a building’s foundation. They occasionally enter buildings through holes or cracks in walls and foundations.
Are Red Ants Harmful to Humans?: Yes! Red imported ants do cause harm. The sting (bite) of a red imported fire ant is very painful and often results in a raised welt that becomes a white blister. People who are allergic to insect stings will react even more severely. Plus, the building of their big mounds can damage plant roots and lead to loss of crops.
What Should I Do If I See Red Imported Fire Ants?: The first thing to do if you see important fire ants is to get away from them. If they start crawling on you, they are likely to bite you, and if there are a bunch of them, that could result in a lot of pain.
Next, immediately ask a grown-up to call a pest control professional. Other types of ants are annoying and can contaminate your food, but at least they can’t physically hurt you. You want these ants away from your house as quickly as possible. There is more info on red imported fire ants and how to get rid of them at the official National Pest Management Association website.
Also watch red imported fire ants face off with Formosan termites on episode 3 of Bug Battles!
More fun with pests
Games
Videos
Crafts
Books
Parents & Teachers:
We don't allow any on-site or pop-up advertising, or third-party external links, so children can play and learn freely!
Content for Grade Levels:
Pre-K Activities
Elementary Grade Activities
Middle School Grade Activities
Brought to you by the National Pest Management Association.
Visit PestWorld.org for additional information on bugs and pest control experts.
Contact PestWorld for Kids
•
© Copyright 2024 National Pest Management Association
<
Ant - Simple English Wikipedia, the free encyclopedia
Jump to content
Main menu
Main menu
move to sidebar
hide
Getting around
Main pageSimple startSimple talkNew changesShow any pageHelpContact usGive to WikipediaAbout Wikipedia
Search
Search
Create account
Log in
Personal tools
Create account Log in
Pages for logged out editors learn more
ContributionsTalk
Contents
move to sidebar
hide
Beginning
1Colonies
2Evolution
3Development and reproduction
Toggle Development and reproduction subsection
3.1Mating
3.1.1The nuptial flight
4Uses
5Related pages
6Notes
7Further reading
8Other websites
Toggle the table of contents
Ant
193 languages
AfrikaansAlemannischአማርኛअंगिकाÆngliscالعربيةAragonésܐܪܡܝܐԱրեւմտահայերէնArmãneashtiঅসমীয়াAsturianuAtikamekwAvañe'ẽAymar aruAzərbaycancaتۆرکجهBasa BaliBamanankanবাংলাBanjarBân-lâm-gúBasa BanyumasanБашҡортсаБеларускаяБеларуская (тарашкевіца)Bikol CentralБългарскиབོད་ཡིགBosanskiBrezhonegБуряадCatalàЧӑвашлаCebuanoČeštinaChiShonaCymraegDagbanliDanskDeutschDiné bizaadEestiΕλληνικάEmiliàn e rumagnòlEnglishEspañolEsperantoEuskaraفارسیFiji HindiFrançaisGaeilgeGàidhligGalegoГӀалгӀайગુજરાતી客家語/Hak-kâ-ngîХальмг한국어Հայերենहिन्दीHrvatskiIdoIlokanoBahasa IndonesiaInterlinguaИронÍslenskaItalianoעבריתJawaKabɩyɛಕನ್ನಡKapampanganქართულიकॉशुर / کٲشُرҚазақшаKernowekKiswahiliКомиKongoKreyòl ayisyenKurdîКыргызчаКырык марыLadinЛаккуLatinaLatviešuLëtzebuergeschЛезгиLietuviųLi NihaLimburgsLivvinkarjalaLombardMagyarМакедонскиMalagasyമലയാളംमराठीმარგალურიمصرىمازِرونیBahasa Melayuꯃꯤꯇꯩ ꯂꯣꯟ閩東語 / Mìng-dĕ̤ng-ngṳ̄Монголမြန်မာဘာသာNa Vosa VakavitiNederlandsNedersaksiesनेपालीनेपाल भाषा日本語НохчийнNordfriiskNorsk bokmålNorsk nynorskOccitanОлык марийଓଡ଼ିଆOʻzbekcha / ўзбекчаਪੰਜਾਬੀPälzischPangcahپنجابیپښتوПерем комиPinayuananPlattdüütschPolskiPortuguêsRipoarischRomânăRumantschRuna SimiРусиньскыйРусскийСаха тылаSakizayaᱥᱟᱱᱛᱟᱲᱤSarduScotsShqipSicilianuසිංහලسنڌيSlovenčinaSlovenščinaŚlůnskiSoomaaligaکوردیСрпски / srpskiSrpskohrvatski / српскохрватскиSundaSuomiSvenskaTagalogதமிழ்TaclḥitTaqbaylitТатарча / tatarçaతెలుగుไทยThuɔŋjäŋТоҷикӣᏣᎳᎩತುಳುTürkçeTürkmençeTyapУдмуртУкраїнськаاردوئۇيغۇرچە / UyghurcheVahcuenghVepsän kel’Tiếng ViệtWalonWayuunaiki文言West-VlamsWinarayWolof吴语ייִדיש粵語ZazakiŽemaitėška中文Batak Toba
Change links
PageTalk
English
ReadChangeChange sourceView history
Tools
Tools
move to sidebar
hide
Actions
ReadChangeChange sourceView history
General
What links hereRelated changesUpload fileSpecial pagesPermanent linkPage informationCite this pageGet shortened URLDownload QR codeWikidata item
Print/export
Make a bookDownload as PDFPage for printing
In other projects
Wikimedia CommonsWikispecies
From Simple English Wikipedia, the free encyclopedia
AntsTemporal range: 100 mya–present.[1] Lower Cretaceous – Present
A group of fire ants.
Scientific classification
Domain:
Eukaryota
Kingdom:
Animalia
Phylum:
Arthropoda
Class:
Insecta
Order:
Hymenoptera
Superfamily:
Formicoidea
Family:
FormicidaeLatreille, 1809
Type species
Lasius nigerLinnaeus, 1758
Cladogram of subfamilies
Martialinae
Leptanillinae
Amblyoponinae
Paraponerinae
Agroecomyrmecinae
Ponerinae
Proceratiinae
Ecitoninae‡
Aenictinae‡
Dorylini‡
Aenictogitoninae‡
Cerapachyinae‡*
Leptanilloidinae‡
Dolichoderinae
Aneuretinae
Pseudomyrmecinae
Myrmeciinae
Ectatomminae
Heteroponerinae
Myrmicinae
Formicinae
A phylogeny of the living ant subfamilies.[2][3]
*Cerapachyinae is paraphyletic
‡ The previous dorylomorph subfamilies were synonymized under Dorylinae by Brady et al. in 2014[4]
An example of symbiosis: the ant protects the aphids and harvests their sugary excretion.
Ants are a kind of insect that lives together in large colonies. They are the family Formicidae.[5][6]
Ants are a lot like bees and wasps. They all came from the same kind of animal a long time ago, but now they are different. There are about 22,000 different kinds of ants, but we have only documented 12,500.[7][8][9] Every kind of ant has a thin middle, called a thorax, and two long rods on their heads called antennae.
Ants live in groups that can be big or small. Some kinds of ants live in small groups and eat other animals. Some ants work together in very big groups. These groups can have millions of ants in them that travel outside every day in a big area. Ants are small, but they are very strong. Some ants are strong enough to carry things that are as heavy as 10 times their own weight. Some ants are called workers. Workers dig tunnels and carry food back to the colony so that other ants and the queen ant can eat.[10] The Queen ant is the mother of all the ants in the colony. She is the only non-worker ant.
Colonies[change | change source]
The groups that ants live in are called colonies. A colony has a female ant called a queen which lays eggs. Those eggs will grow into more ants. Big colonies of ants have different kinds of ants that grow from the eggs. These are called different castes of ants. Some are workers which do jobs like carrying and digging, and soldiers which fight other animals. Worker and soldier ants are females. Another type of ant are drones which are male ants.[11]
Really big ant colonies are sometimes called superorganisms. This means the ants work together so well that they are like little parts of one big animal. Ants cannot live by themselves for very long because they need to work with other ants.[12][13]
Ants have colonies almost everywhere on planet Earth. Antarctica lacks ants because it's very cold and there's not much food. Small islands may not have ants.
Evolution[change | change source]
Ants fossilised in Baltic amber
The family Formicidae belongs to the order Hymenoptera, which also includes bees, sawflies and wasps. Ants evolved from a lineage within the vespoid wasps.
Phylogenetic analysis suggests that ants arose in the Lower Cretaceous period about 110 to 130 million years ago, or even earlier. One estimate from DNA studies places the origin of ants at ≈140 million years ago (mya).[14] Another study puts it in the Jurassic at 185 ± 36 mya (95% confidence limits).[15]
After the rise of flowering plants about 100 million years ago ants diversified. They became ecologically dominant about 60 million years ago.[16][17][18]
In 1966 E.O. Wilson and his colleagues identified the fossil remains of an ant (Sphecomyrma freyi) from the Cretaceous period. The specimen, trapped in amber dating back to more than 80 million years ago, has features of both ants and wasps.[19] Sphecomyrma was probably a ground forager but some suggest that primitive ants were likely to have been predators underneath the surface of the soil.
During the Cretaceous period, a few species of primitive ants ranged widely on the Laurasian super-continent (the northern hemisphere). They were scarce in comparison to other insects, representing about 1% of the insect population.
Ants became dominant after adaptive radiation at the beginning of the Cainozoic. By the Oligocene and Miocene ants had come to represent 20-40% of all insects found in major fossil deposits. Of the species that lived in the Eocene epoch, approximately one in ten genera survive to the present. Genera surviving today comprise 56% of the genera in Baltic amber fossils (early Oligocene), and 92% of the genera in Dominican amber fossils (apparently early Miocene).[16][20]p23
Termites, though sometimes called white ants, are not ants and belong to the order Isoptera. Termites are actually more closely related to cockroaches and mantids. Termites are eusocial but differ greatly in the genetics of reproduction. The similar social structure is attributed to convergent evolution.[21] Velvet ants look like large ants, but are wingless female wasps.[22][23]
Development and reproduction[change | change source]
Meat eater ant nest during swarming.
The life of an ant starts from an egg. If the egg is fertilised, the progeny will be female (diploid); if not, it will be male (haploid). Ants develop by complete metamorphosis with the larval stages passing through a pupal stage before emerging as an adult. The larva is fed and cared for by workers.
Food is given to the larvae by trophallaxis, a process in which an ant regurgitates liquid food held in its crop. This is also how adults share food, stored in the 'social stomach', among themselves.
Larvae may also be given solid food brought back by foraging workers, and may even be taken to captured prey in some species. The larvae grow through a series of moults and enter the pupal stage.[24]
The differentiation into queens and workers (which are both female), and different castes of workers, is influenced in some species by the food the larvae get. Genetic influences, and the control of gene expression by the feeding are complex. The determination of caste is a major subject of research.[20]p351, 372[25]
A new worker spends the first few days of its adult life caring for the queen and young. It then does digging and other nest work, and later, defends the nest and forages. These changes are sometimes fairly sudden, and define what are called temporal castes. An explanation for the sequence is suggested by the high casualties involved in foraging, making it an acceptable risk only for ants that are older and are likely to die soon of natural causes.[26][27]
Mating[change | change source]
Most ant species have a system in which only the queen and breeding females can mate. Contrary to popular belief, some ant nests have multiple queens (polygyny). The life history of Harpegnathos saltator is
exceptional among ants because both queens and some workers
reproduce sexually.[28]
The winged male ants, called drones, emerge from pupae with the breeding females (although some species, like army ants, have wingless queens), and do nothing in life except eat and mate.
The nuptial flight[change | change source]
Most ants produce a new generation each year.[29] During the species specific breeding period, new reproductives, winged males and females leave the colony in what is called a nuptial flight. Typically, the males take flight before the females. Males then use visual cues to find a common mating ground, for example, a landmark such as a pine tree to which other males in the area converge. Males secrete a mating pheromone that females follow. Females of some species mate with just one male, but in some others they may mate with anywhere from one to ten or more different males.[20] Mated females then seek a suitable place to begin a colony. There, they break off their wings and begin to lay and care for eggs. The females store the sperm they obtain during their nuptial flight to selectively fertilise future eggs.
The first workers to hatch are weak and smaller than later workers, but they begin to serve the colony immediately. They enlarge the nest, forage for food and care for the other eggs. This is how new colonies start in most species. Species that have multiple queens may have a queen leaving the nest along with some workers to found a colony at a new site,[20]p143 a process akin to swarming in honeybees.
Ants mating.
A wide range of reproductive strategies have been noted in ant species. Females of many species are known to be capable of reproducing asexually through parthenogenesis,[30] and one species, Mycocepurus smithii is known to be all-female.[31]
Ant colonies can be long-lived. The queens can live for up to 30 years, and workers live from 1 to 3 years. Males, however, are more transitory, and survive only a few weeks.[32] Ant queens are estimated to live 100 times longer than solitary insects of a similar size.[33]
Ants are active all year long in the tropics but, in cooler regions, survive the winter in a state of dormancy or inactivity. The forms of inactivity are varied and some temperate species have larvae going into the inactive state (diapause), while in others, the adults alone pass the winter in a state of reduced activity.[34]
Uses[change | change source]
It may seem strange that ants have uses, but there are some. Some people use ants for food, medicine and rituals. Some species of ants are used for pest control (they eat pests that destroy food for humans). They can damage crops and enter buildings, though. Some species, like the red imported fire ant, live in places where they came to by complete accident.
Related pages[change | change source]
Formic acid
Leaf-cutter ant
Notes[change | change source]
↑ Moreau CS, Bell CD, Vila R, Archibald SB, Pierce NE (April 2006). "Phylogeny of the ants: diversification in the age of angiosperms". Science. 312 (5770): 101–4. Bibcode:2006Sci...312..101M. doi:10.1126/science.1124891. PMID 16601190. S2CID 20729380.
↑ Ward, Philip S (2007). "Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae)" (PDF). Zootaxa. 1668: 549–563. doi:10.11646/zootaxa.1668.1.26.
↑ Rabeling C, Brown JM, Verhaagh M (2008). "Newly discovered sister lineage sheds light on early ant evolution". PNAS. 105 (39): 14913–7. Bibcode:2008PNAS..10514913R. doi:10.1073/pnas.0806187105. PMC 2567467. PMID 18794530.
↑ Brady, Seán G; Fisher, Brian L; Schultz, Ted R; Ward, Philip S (2014). "The rise of army ants and their relatives: diversification of specialized predatory doryline ants". BMC Evolutionary Biology. 14: 2–14. doi:10.1186/1471-2148-14-93. PMC 4021219. PMID 24886136.
↑ pronounced /fɔrˈmɪsɨdiː/). The word ant comes from ante, a Middle English which comes from æmette of Old English and is related to the Old High German āmeiza. All of these words come from West Germanic *amaitjo. It meant "the biter" (from *ai-, "off, away" and *mait- "cut"). ""ant". Merriam-Webster Online Dictionary". Merriam-Webster. Retrieved 6 June 2008.
↑ "Ant. Online Etymology Dictionary". Retrieved 30 May 2009. The family name Formicidae is from the Latin word formīca ("ant"). Simpson DP (1979). Cassell's Latin Dictionary (5 ed.). London: Cassell Ltd. ISBN 978-0-304-52257-6. The word "ant" in other languages such as the Portuguese formiga, Italian formica, Spanish hormiga, Romanian furnică and French fourmi come from the Latin word.
↑ Bolton, Barry (1995). A new General Catalogue of the Ants of the World. Harvard University Press. ISBN 978-0-674-61514-4.
↑ "Hymenoptera name server. Formicidae species count". Ohio State University. Archived from the original on 2008-06-18. Retrieved 2010-12-06.
↑ La nueva taxonomía de hormigas. Pages 45-48 in Fernández, F. Introducción a las hormigas de la región neotropical (PDF). Instituto Humboldt, Bogotá. 2003. {{cite book}}: Cite uses deprecated parameter |authors= (help)
↑ Wade, Nicholas (15 July 2008). "Taking a cue from ants on evolution of humans". New York Times. Retrieved 15 July 2008.
↑ Oster, George F.; Wilson, Edward O. (1978). Caste and ecology in the social insects. Vol. 12. Princeton University Press, Princeton. pp. 21–22. ISBN 978-0-691-02361-8. PMID 740003. {{cite book}}: |journal= ignored (help)
↑ Oster, George F.; Wilson, Edward O. (1978). Caste and ecology in the social insects. Vol. 12. Princeton University Press, Princeton. pp. 21–22. ISBN 978-0-691-02361-8. PMID 740003. {{cite book}}: |journal= ignored (help)
↑ Flannery, Tim (2011). A Natural History of the Planet. Grove/Atlantic, Inc. p. 79. ISBN 978-0-8021-9560-9.
↑ Brady S.G. 2003. Evolution of the army ant syndrome: the origin and long-term evolutionary stasis of a complex of behavioral and reproductive adaptations. Proc. Natl. Acad. Sci. USA. 100, 6575–6579.Evolution of the army ant syndrome: The origin and long-term evolutionary stasis of a complex of behavioral and reproductive adaptations | PNAS
↑ Crozier R.H, Jermiin L.S. & Chiotis M. 1997. Molecular evidence for a Jurassic origin of ants. Naturwissenschaften 84, 22–23.
↑ 16.0 16.1 Grimaldi D, Agosti D (2001). "A formicine in New Jersey Cretaceous amber (Hymenoptera: Formicidae) and early evolution of the ants". Proceedings of the National Academy of Sciences. 97 (25): 13678–13683. doi:10.1073/pnas.240452097. PMC 17635. PMID 11078527.
↑ Moreau CS; et al. (2006). "Phylogeny of the ants: diversification in the Age of Angiosperms". Science. 312 (5770): 101–104. Bibcode:2006Sci...312..101M. doi:10.1126/science.1124891. PMID 16601190. S2CID 20729380.
↑ Wilson E.O, Hölldobler B (2005). "The rise of the ants: A phylogenetic and ecological explanation". Proceedings of the National Academy of Sciences. 102 (21): 7411–7414. Bibcode:2005PNAS..102.7411W. doi:10.1073/pnas.0502264102. PMC 1140440. PMID 15899976.
↑ Wilson E.O., Carpenter FM, Brown WL (1967). "The first Mesozoic ants". Science. 157 (3792): 1038–1040. Bibcode:1967Sci...157.1038W. doi:10.1126/science.157.3792.1038. PMID 17770424. S2CID 43155424.{{cite journal}}: CS1 maint: multiple names: authors list (link)
↑ 20.0 20.1 20.2 20.3 Holldobler, Bert; Wilson, Edward O. (1998). The Ants. Springer. ISBN 978-3-540-52092-4.
↑ Thorne, Barbara L (1997). "Evolution of eusociality in termites" (PDF). Annu. Rev. Ecol. Syst. 28 (5): 27–53. doi:10.1146/annurev.ecolsys.28.1.27. PMC 349550. Archived from the original (PDF) on 2010-05-30. Retrieved 2010-12-07.
↑ "Order Isoptera - Termites". Iowa State University Entomology. 2004. Retrieved 12 June 2008.
↑ "Family Mutillidae - Velvet ants". Iowa State University Entomology. 2004. Retrieved 12 June 2008.
↑ Gillott, Cedric (1995). Entomology. Springer. p. 325. ISBN 978-0-306-44967-3.
↑ Anderson, Kirk E.; Linksvayer, Timothy A.; Smith, Chris R. (2008). "The causes and consequences of genetic caste determination in ants (Hymenoptera: Formicidae)" (PDF). Myrmecol. News. 11: 119–132. Archived from the original (PDF) on 2011-07-27. Retrieved 2010-12-07.
↑ Traniello JFA (1989). "Foraging strategies of ants". Annual Review of Entomology. 34: 191–210. doi:10.1146/annurev.en.34.010189.001203.
↑ Sorensen A, Busch TM, Vinson SB (1984). "Behavioral flexibility of temporal sub-castes in the fire ant, Solenopsis invicta, in response to food". Psyche. 91 (3–4): 319–332. doi:10.1155/1984/39236.{{cite journal}}: CS1 maint: multiple names: authors list (link)
↑ Peeters C, Holldobler B (1995). "Reproductive cooperation between queens and their mated workers: The complex life history of an ant with a valuable nest" (PDF). Proceedings of the National Academy of Sciences. 92 (24): 10977–10979. Bibcode:1995PNAS...9210977P. doi:10.1073/pnas.92.24.10977. PMC 40553. PMID 11607589.
↑ Taylor, Richard W. (2007). "Bloody funny wasps! Speculations on the evolution of eusociality in ants". In Snelling R.R., B.L. Fisher, & P S. Ward (ed.). Advances in ant systematics (Hymenoptera: Formicidae): homage to E.O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute, 80. American Entomological Institute. pp. 580–609.{{cite book}}: CS1 maint: multiple names: editors list (link)
↑ Heinze, Jurgen; Tsuji, Kazuki (1995). "Ant reproductive strategies" (PDF). Res. Popul. Ecol. 37 (2): 135–149. doi:10.1007/BF02515814. S2CID 21948488. Archived from the original (PDF) on 2011-05-27. Retrieved 2010-12-07.
↑ Himler, Anna G.; Caldera, EJ; Baer, BC; Fernández-Marín, H; Mueller, UG (2009). "No sex in fungus-farming ants or their crops". Proc. R. Soc. B. 276 (1667): 2611–2616. doi:10.1098/rspb.2009.0313. PMC 2686657. PMID 19369264.
↑ Keller L (1998). "Queen lifespan and colony characteristics in ants and termites". Insectes Sociaux. 45 (3): 235–246. doi:10.1007/s000400050084. S2CID 24541087.
↑ Resh, Vincent H.; Carde, Ring T. (2003). Encyclopedia of Insects. San Diego: Academic Press. pp. 29–32. ISBN 978-0-12-586990-4.
↑ Kipyatkov V.E (2001). "Seasonal life cycles and the forms of dormancy in ants (Hymenoptera, Formicoidea)". Acta Societatis Zoologicae Bohemicae. 65 (2): 198–217.
Further reading[change | change source]
Borror, Donald Joyce; Johnson, Norman F.; Triplehorn, Charles A. (1989). An introduction to the study of insects. Brooks/Cole Publishing Company.
Hölldobler B. and Wilson E.O. 1998. Journey to the ants: a story of scientific exploration. Harvard University Press. ISBN 978-0-674-48526-6
Holldobler, Bert; Wilson, Edward O. (1998). The Ants. Springer. ISBN 978-3-540-52092-4.
Holldobler, Bert; Wilson, Edward O. (2009). The superorganism: the beauty, elegance, and strangeness of insect societies. W.W. Norton. ISBN 978-0-393-06704-0.
Other websites[change | change source]
Wikispecies has information on: ants.
Media related to Formicidae at Wikimedia Commons
"Antweb from The California Academy of Sciences". antweb.org. Retrieved 6 December 2010.
"AntBlog, a website for studying ant colonies". antblog.co.uk. Archived from the original on 19 November 2016. Retrieved 6 December 2010.
Global Ant Project Archived 2009-02-12 at the Wayback Machine
The super-nettles. A dermatologist's guide to ants-in-the-plants
Ant Citizendium
Retrieved from "https://simple.wikipedia.org/w/index.php?title=Ant&oldid=9266971"
Category: AntsHidden categories: Articles containing Middle English (1100-1500)-language textArticles containing Old English (ca. 450-1100)-language textArticles containing Old High German (ca. 750-1050)-language textArticles with text in West Germanic languagesArticles with text in Germanic languagesArticles containing Latin-language textArticles containing Portuguese-language textArticles containing Italian-language textArticles containing Spanish-language textArticles containing Romanian-language textArticles containing French-language textCS1 errors: deprecated parametersCS1 errors: periodical ignoredCS1 maint: multiple names: authors listCS1 maint: multiple names: editors listArticles with 'species' microformatsTaxobox articles missing a taxonbarCommons category link is on WikidataWebarchive template wayback links
This page was last changed on 26 December 2023, at 00:57.
Text is available under the Creative Commons Attribution-ShareAlike License and the GFDL; additional terms may apply. See Terms of Use for details.
Privacy policy
About Wikipedia
Disclaimers
Code of Conduct
Developers
Statistics
Cookie statement
Mobile view
Toggle limited content width
Ant: characteristics, diet, and colony structure | Britannica
Search Britannica
Click here to search
Search Britannica
Click here to search
Login
Subscribe
Subscribe
Home
Games & Quizzes
History & Society
Science & Tech
Biographies
Animals & Nature
Geography & Travel
Arts & Culture
Money
Videos
On This Day
One Good Fact
Dictionary
New Articles
History & Society
Lifestyles & Social Issues
Philosophy & Religion
Politics, Law & Government
World History
Science & Tech
Health & Medicine
Science
Technology
Biographies
Browse Biographies
Animals & Nature
Birds, Reptiles & Other Vertebrates
Bugs, Mollusks & Other Invertebrates
Environment
Fossils & Geologic Time
Mammals
Plants
Geography & Travel
Geography & Travel
Arts & Culture
Entertainment & Pop Culture
Literature
Sports & Recreation
Visual Arts
Companions
Demystified
Image Galleries
Infographics
Lists
Podcasts
Spotlights
Summaries
The Forum
Top Questions
#WTFact
100 Women
Britannica Kids
Saving Earth
Space Next 50
Student Center
Home
Games & Quizzes
History & Society
Science & Tech
Biographies
Animals & Nature
Geography & Travel
Arts & Culture
Money
Videos
ant
Table of Contents
ant
Related Summaries
E.O. Wilson Summary
Discover
10 Questions About Insects Answered
8 Must-See Paintings at the National Gallery of Art in Washington, D.C.
Celebrating Ramadan
How Did Helen Keller Fly a Plane?
How Did Alexander the Great Really Die?
Titanosaurs: 8 of the World's Biggest Dinosaurs
7 Famous Child Prodigies
Home
Science
Bugs, Mollusks & Other Invertebrates
Insects
ant Article
ant summary
Actions
Cite
verifiedCite
While every effort has been made to follow citation style rules, there may be some discrepancies.
Please refer to the appropriate style manual or other sources if you have any questions.
Select Citation Style
MLA
APA
Chicago Manual of Style
Copy Citation
Share
Share
Share to social media
URL
https://www.britannica.com/summary/ant
Know about ants, their characteristics, diet, and colony structure
Cite
verifiedCite
While every effort has been made to follow citation style rules, there may be some discrepancies.
Please refer to the appropriate style manual or other sources if you have any questions.
Select Citation Style
MLA
APA
Chicago Manual of Style
Copy Citation
Share
Share
Share to social media
URL
https://www.britannica.com/summary/ant
Written and fact-checked by
The Editors of Encyclopaedia Britannica
Encyclopaedia Britannica's editors oversee subject areas in which they have extensive knowledge, whether from years of experience gained by working on that content or via study for an advanced degree. They write new content and verify and edit content received from contributors.
The Editors of Encyclopaedia Britannica
Below is the article summary. For the full article, see ant.
Carpenter ant (Camponotus).ant, Any member of approximately 10,000 species of the social insect family Formicidae. Ants are found worldwide but are especially common in hot climates. They range from 0.1 to 1 in. (2–25 mm) long and are usually yellow, brown, red, or black. Ants eat both plant and animal substances; some even “farm” fungi for food, cultivating them in their nests, or “milk” aphids. Ant colonies consist of three castes (queens, males, and workers, including soldiers) interacting in a highly complex society paralleling that of the honeybees. Well-known ant species are the carpenter ants of North America, the voracious army ants of tropical America, and the stinging fire ant.
E.O. Wilson Summary
E.O. Wilson American biologist recognized as the world’s leading authority on ants. He was also the foremost proponent of sociobiology, the study of the genetic basis of the social behaviour of all animals, including humans. (Read E.O. Wilson’s Britannica essay on mass extinction.) Wilson received
ant
- Students | Britannica Kids | Homework Help
Articles
Animals
Arts and Literature
Earth and Geography
History
Life Processes
Living Things (Other)
Philosophy and Religion
Plants
Science and Mathematics
Society
Sports and Recreation
Technology
Images & Video
Animals
Arts and Literature
Earth and Geography
History
Life Processes
Living Things (Other)
Philosophy and Religion
Plants
Science and Mathematics
Society
Sports and Recreation
Technology
Biographies
Dictionary
Compare Countries
World Atlas
Podcast
Switch Level
Kids
Students
Scholars
About Us
students
Kids
Students
Scholars
Fundamentals
NEW
7-day free trial
Search
Login
Articles
Featured Article
national park
All Categories
Animals
Arts and Literature
Earth and Geography
History
Life Processes
Living Things (Other)
Philosophy and Religion
Plants
Science and Mathematics
Society
Sports and Recreation
Technology
Images & Videos
Featured Media
George Washington and the Continental Congress
All Categories
Animals
Arts and Literature
Earth and Geography
History
Life Processes
Living Things (Other)
Philosophy and Religion
Plants
Science and Mathematics
Society
Sports and Recreation
Technology
Biographies
Dictionary
Compare Countries
World Atlas
Podcast
×
ant
View article for:
Kids
Students
Scholars
Article
Images & Videos
Related
Subscriber features
Print (Subscriber Feature)
Email (Subscriber Feature)
Cite (Subscriber Feature)
Translate (Subscriber Feature)
Listen (Subscriber Feature)
×
Contents
IntroductionDistribution and HabitatPhysical CharacteristicsBehaviorLife CycleTypes of AntsRelationship to Humans
Did You Know?
A swarm of army ants can destroy all crops and any small animals in its path.
Related resources for this article
Articles
Primary Sources & E-Books
Websites
View search results for:
Search
Introduction
AdstockRFEncyclopædia Britannica, Inc.
Ants are small insects that are found throughout the world. They are known for carrying objects much heavier and bigger than themselves. Ants are social insects. They live in organized communities, work cooperatively and efficiently, create a clear division of labor, wage war, and occasionally capture slaves. Ants belong to the order Hymenoptera, which also includes bees and wasps. Ants constitute the family Formicidae, which has more than10,000 species. Entomologists, or scientists who study insects, have estimated that there are probably more ants than any other kind of insect. Ants have been in existence for more than 100 million years, and they probably evolved from wasps. Distribution and Habitat© Ljupco Smokovski/Fotolia
Ants occur throughout the world but are particularly abundant in hot climates. They have traveled from their original home, the tropics, to areas as varied as the polar regions, mountain ranges, and deserts. They make their nests in many materials, including soil, sand, wood, and leaves.Physical CharacteristicsDespite their great diversity in social behavior and habits, most ants have the same basic physical structure. They range in size from 0.08 to 1 inch (2 to 25 millimeters). They are usually yellow, brown, red, or black. A few have a metallic luster.Encyclopædia Britannica, Inc.
The bodies of ants, like those of all insects, are divided into three sections: the head, the thorax, and the abdomen. The head is large, and the abdomen is slender and oval. The six legs are attached to the thorax, or midsection. The thorax is connected to the abdomen by a small “waist” section. The ant’s vital organs are located in the abdomen.The mouth is an important working tool for most ants. It consists of two sets of jaws. The outer pair is used for carrying objects such as food and for digging. The inner pair is used for chewing.Adult ants swallow only liquids. In the back of the jaw is a storage pocket. Solid food goes first into this pocket, where a strong saliva breaks it down. The ant swallows the liquid part. The solids form into a ball that the ant spits out.© Morley Read/Dreamstime.com
Different parts of ants vary according to the way a particular species lives. Harvester ants have short, heavy, crushing jaws for breaking seeds. Leaf-cutters have jaws with saw-toothed edges so that they can shred leaves. The jaws of the amazon ant are sickle-shaped, like a curved tool. They use these jaws to kill other insects efficiently.Most ants have simple eyes, called ocelli, on top of the head as well as a compound eye with many lenses on each side of the head. Nonetheless, most species have poor vision. More useful than the eyes are the antennae, which serve as the main sensory organs. The antennae are the two slender, pointed rods that wave constantly from the head as the ant moves about. Each antenna has a joint at the midpoint, enabling it to bend like an elbow.Walter Dawn
The senses of smell and touch are located in the antennae. With these rods, the ant recognizes its nest and the members of its colony. It can instantly detect an intruder from another colony by its smell. Ants also are able to communicate with one another by tapping with their antennae. Nursemaid ants clean the young with their antennae as well as with their tongues. Ants also clean themselves with a sort of comb called a strigil, which is located on the forelegs.Males and queens are normally winged, although they use these wings only once—on their mating flight. All other ants lack wings.K. Fogassy/B.W. Halstead, World Life Research Institute
Many ants have only simple stings, and instead of stinging their enemies they eject vapors of formic acid. Other ants, such as bulldog ants and fire ants, have powerful stings at the tip of the abdomen.Ants do not have lungs. They breathe instead through small holes called spiracles, which are located along the sides of the abdomen and the thorax.Behavior© Forest71/Dreamstime.comEncyclopædia Britannica, Inc.
The most distinguishing trait of
ant behavior is sociability. Ants do not act individually. Instead, they behave according to the needs of the colony in roles dictated by the caste—or job class, such as worker or soldier—into which they are born. The major social unit is the colony, which forms a nest.AdstockRF
The nests of ants vary in structure and in material. Most of an ant’s life is spent in its nest. The number of ants living in one nest ranges from a few to more than a million. Some ants dig chambers and passages in the ground. Others locate their nests under rocks, in trees, or in logs. The nests may be built of paper, twigs, sand, gravel, or other materials.One colony can claim as its territory thousands of square yards (more than a thousand square meters). Some species pile large heaps of earth on top of their nests. These mounds can be more than 1 yard (0.9 meter) high and up to 4 yards (3.7 meters) in diameter. The nest is kept spotless. The ants remove all rubbish promptly. If an ant dies, a worker carries it from the nest to a spot that serves as both a cemetery and a garbage dump. If the queen dies, however, the workers continue to care for her as long as her body remains recognizable.Ants like warmth and swarm out of their nests into the sunshine. But they can be frozen for long periods without harm. Packed in tight bunches, many spend the winter inside logs and stumps. Others lie under plant roots or in shallow nests in the ground. Certain species deepen their nests below the frost line as winter approaches. There they crowd into their tunnels and chambers with their legs interlocked and sleep through the cold winter.The social structure of the colony has led to several unusual living arrangements. Ants can be very hospitable. Some species live as guests with other ants or insects, and some ants host other insects as guests. A few species live with parasites—insects or other small animals, such as mites, spiders, caterpillars, and beetles, that are fed and sheltered by the colony but that provide nothing in return.AdstockRF
The sociability of ants is especially evident in their feeding habits. The abdomen has two stomachs. The ant digests part of the food for its own use in the true stomach. The rest goes into a sac called the social stomach, or crop. This is a sort of storage tank for the use of the entire colony.When an ant is hungry, it strokes a worker with its antennae. The worker brings up a drop of liquid from its crop and passes it into the other’s mouth. Thus the queen and other ants are fed by those whose job it is to find and bring back food.Encyclopædia Britannica, Inc.Hospitality and cooperation
are only one aspect of the ant’s social character. Ants sometimes have conflict, either with members of their own species or with members of other species. Because ants are so plentiful, it is common for different species to come into contact. If one side is threatened, such as if another colony encroaches on its territory, then an ant warfare can break out. Ant warfare is usually an organized, aggressive action. Ants use physical attacks, such as biting or stinging. Some are able to release chemicals that confuse or subdue their enemies. Ants often work together in holding an enemy down and destroying it. They also go after the eggs and larvae of their enemies and then raise them as slaves.The dependence on slaves is clearly evident in amazon ants. Because of the shape of their scissorlike jaws, they can easily kill prey and capture the pupae of other ant species. However, amazon ants are incapable of other types of work, including feeding themselves. They use the captured species in their colony to care for their young, expand the nest, and do the other chores of the worker caste.The ant’s aggressive nature is also seen in robberies and quarrels over boundaries, which can lead to feuds lasting for years between different colonies of ants. Ants vary in their methods of defense. Some bite or spit out a disagreeable liquid. Others use their stings to shoot out formic acid. Still others run away when under attack or “play dead.” Some ants rub their legs along their abdomen to make sound signals to warn other members of their colonies.Scott Bauer—ARS/USDA
In addition to being able to communicate, ants have an excellent sense of direction. They can find their way back to their nest by vision and smell. They orient themselves by the position of the Sun and by memory of landmarks, such as trees. Some ants also leave scent trails to aid other ants.Life Cycle© Tomatito26 /Dreamstime.com
The life cycle of an ant generally has four stages: egg, larva, pupa, and adult. Most ants live from 8 to 10 weeks, although certain queens may live for as long as 15 years, and some workers for up to 7 years.The role, or caste, is the occupational group into which an individual ant is born. There are three major castes of ants: queens, workers, and soldiers. The queen is the mother and founder of the colony. A colony may have one queen or many queens, depending on the ant species. The queen spends her life laying eggs. The workers are sterile females and do the jobs necessary to keep the colony in good working order. They care for the young, enlarge the nest, and gather food to feed the queen and the other members of the colony. The larger workers—the soldiers—defend the nest. They raid other colonies and often capture slaves.At certain times of the year, many species produce winged males and queens, which fly into the air, where they mate. The male deposits sperm in the queen’s sperm sac. The winged males die soon after mating with the queens. They are not allowed to return to their original nests, so they starve or are eaten by other insects or by birds.The queen digs a hole where she lays tiny, white eggs. As she lays them she may or may not fertilize them with sperm from her sperm sac. Fertilized eggs result in females, and unfertilized eggs produce males. The female eggs develop into fertile queens, sterile workers, or soldiers. Most of the ants in a colony are workers.While the queen waits for her eggs to hatch, she is nourished by food stored in her body. The first eggs usually produce workers, which the queen feeds with other eggs. When the workers are able to bring in food from outside the nest, the queen devotes the rest of her life to laying eggs.AdstockRF
Ant eggs develop into larvae in a week or two. The larvae are white and oblong, and they lack eyes and legs. Unlike adult ants, they can eat solid food, which the earliest workers obtain for them. The larvae eat constantly and continually grow, going through several molts in which they shed their outer skins. The nurses lick the larvae constantly, in what looks like a show of affection. Actually, the nurse workers are eating a sweet liquid that appears on the larvae.After about a month the larvae change into pupae. This is a last resting stage before the ants emerge. Pupae resemble adult ants except that they are still whitish in color and the legs and antennae are folded in toward the body. Some pupae spin a silken cocoon about their bodies and rest inside, but others remain naked. It takes about 6 to 10 weeks for the pupae to turn into adults. The newly hatched adults are usually lighter and softer than older adults. However, they darken and their exoskeletons harden over time.Types of AntsAlthough ants share many physical and social traits, there are many distinct varieties that differ in their habits and appearance. They are classified according to the shape of certain parts of their bodies and by specific behaviors.Scientists divide ants into several subfamilies, including Dorylinae, Formicinae, and Myrmicinae. Dorylines are predatory ants living in the tropics and many temperate areas. There are about 680 described species.Formicines are found throughout the world. There are some 3,200 living species. They possess venom glands, with which they spray or drip formic acid on their enemies to stun or kill them. Myrmicinae is the most widespread subfamily. This is reflected in the word myrmecology, which means the scientific study of ants. There are some 7,000 known species of myrmicines. Most have a stinger and a sound-producing organ.E.S. Ross
Army ants (subfamily Dorylinae) are found in the tropics. They are sometimes called driver ants or legionary ants. There are about 200 species of army ants. They are nomadic and are notorious for the destruction of plant and animal life in their path. These wanderers do not build permanent nests; instead, they cling together on logs or in hollow trees. They travel in columns, sometimes at speeds of up to 65 feet (20 meters) per hour. Some of these columns have been said to contain up to 20 million ants. Periodically, the colony rests for several days while the queen lays her eggs. As the colony travels, the growing larvae are carried along by the workers. When a swarm of army ants marches through a human settlement, it can destroy all crops and any small animals in its path.Encyclopædia Britannica, Inc.
Leaf-cutter ants (subfamily Myrmicinae) live in the American tropics. There are about 40 species. These ants carry cut-up flowers and leaves overhead like umbrellas, which earns them the nickname of parasol ants. Their foraging columns may be composed of hundreds or thousands of ants. Their trails often stretch more than 100 feet (30 meters) across the forest floor and up and down the trunks of canopy trees. Leaf-cutter ants grow a fungus crop on chewed leaf pieces, carefully tending it like farmers and harvesting it for food.Bulldog, or bull, ants (subfamily Myrmeciinae) are aggressive stinging insects of Australia. There are about 90 species. They are among the largest ants, often reaching 1.6 inches (4 centimeters) in length. They capture insects, which they feed to their young, by grabbing them with their strong mandibles (jaws) and repeatedly stinging them and injecting venom. The adults feed mainly on plant juices.Fire ants (genus Solenopsis) are also called thief ants. They occur in tropical regions of the world, such as in Central and South America, and in some temperate regions, such as in North America. They are red or yellowish and can inflict a severe sting. The nest consists of a loose mound with open craters for ventilation. The workers are notorious for damaging planted grain and attacking poultry. Fire ants communicate through chemical secretions and sounds produced by rubbing or drumming one body part against another. Adult workers are known for their aggressive behavior when under the threat of attack from neighboring ant colonies. Young fire ants, whose stingers and external skeletons are not yet fully developed, play dead.Sahara desert ants (genus Cataglyphis) dwell in the Sahara. They are well adapted to the extreme conditions of their habitat and can tolerate very hot temperatures. Long legs allow them to move rapidly and to elevate their bodies above the scorched sand as they forage for dead insects. These ants are able to return to their nests in a straight line after typically circuitous searches for food up to 330 feet (100 meters) away.© Antagain—iStock/Getty Images
Amazon ants (genus Polyergus) are unable to build nests or feed their young by themselves. They invade the nests of other species, killing the workers and bringing home the helpless young ants to raise as slaves and to do the work of the amazon colony.Encyclopædia Britannica, Inc.Grace Thompson—The National Aubudon Society Collection/Photo Researchers
Many other varieties of ants exist. Carpenter ants (genus Camponotus) live in wood. Harvester ants (genera Messor and Pogonomyrmex) store seeds, grass, and berries as food. Weaver ants (genus Oecophylla) make nests from leaves and other materials held together by silk secreted by the larvae. Honey ants (in various genera) feed on honeydew (a by-product of digestion) that tiny insects called aphids secrete. The ant usually obtains the liquid by gently stroking the aphid’s abdomen with its antennae.Relationship to HumansContunico © ZDF Studios GmbH, Mainz
Although ants are generally considered an annoyance in houses, only a few species are capable of doing real damage to human property. Some species are actually quite helpful to humans. Certain species help get rid of pests. Others, such as ground-dwellers, are useful to farmers. In the process of building their nests, they turn over the soil, which is good for crops. Ants also disperse seeds and pollinate plants.
×
It’s here: the NEW Britannica Kids website!
We’ve been busy, working hard to bring you new features and an updated design. We hope you and your family enjoy the NEW Britannica Kids. Take a minute to check out all the enhancements!
The same safe and trusted content for explorers of all ages.
Accessible across all of today's devices: phones, tablets, and desktops.
Improved homework resources designed to support a variety of curriculum subjects and standards.
A new, third level of content, designed specially to meet the advanced needs of the sophisticated scholar.
And so much more!
Want to see it in action?
Take a tour
Start a free trial
Subscribe now!
×
To
Recipients
Please enter a valid email address.
To share with more than one person, separate addresses with a comma
From
Sender Name
Please enter your name.
Sender Email
Please enter a valid email address.
Cancel
Submit
Translate this page
Choose a language from the menu above to view a computer-translated version of this page. Please note: Text within images is not translated, some features may not work properly after translation, and the translation may not accurately convey the intended meaning. Britannica does not review the converted text.
After translating an article, all tools except font up/font down will be disabled. To re-enable the tools or to convert back to English, click "view original" on the Google Translate toolbar.
About Us
Contact Us
Privacy Notice
Terms of Use
Diversity
©2024 Encyclopædia Britannica, Inc.
This website uses cookies to help deliver and improve our services and provide you with a much richer experience during your visit. To learn more about cookies and your cookie choices, click here.
×